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Male Northern cardinal (Cardinalis cardinalis), in breeding plumage, perched in snow-covered Eastern redbud (Cercis canadensis) tree during spring snowstorm, Freeville, New York, USA. April
Female Polar bear (Ursus maritimus) with cub in snow, Churchill, Canada. November
Red fox (Vulpes vulpes) vixen walking through snow, Derbyshire, UK. January
Mandarin duck (Aix galericulata) drake walking on snow-covered, frozen lake, London, UK. February
Great spotted woodpecker (Dendrocopos major) perched on a snow-covered branch, Germany. February
Goldfinch (Carduelis carduelis) perched on Teasel (Dipsacus sp. ) seedhead in winter, Germany. JanuaryGoldfinch (Carduelis carduelis) perched on Teasel (Dipsacus sp.) seedhead in winter, Germany. January
A female kingfisher (Alcedo atthis) perched on a branch in snow. Leeds, Yorkshire, UK. January
African grey hornbill (Lophoceros nasutus) perched on branch, feeding on large beetle, Tarangire National Park, Tanzania
Toco toucan (Ramphastos toco) perched on branch, holding egg in its bill before swallowing it, Pantanal, Brazil
Pair of Wilson's birds of paradise (Cicinnurus respublica), male performing courtship display to female from a small sapling in the middle of his display court, Batanta Island, West Papua
Vogelkop bowerbird (Amblyornis inornatus) male, at his bower surrounded by orange leaves and petals and other objects he has collected, Arfak Mountains, Vogelkop Peninsula, West Papua
Hoatzin (Opisthocomus hoazin) perched on branch, flapping its wings, above Anangu creek, Yasuni National Park, Francisco de Orellana Province, Ecuador
Ptarmigan (Lagopus mutus) winter plumage, in flight, Utsjoki, Finland. February
Lion (Panthera leo) cub, aged 3 months, climbing on branch and chewing on a stick, Okavango Delta, Botswana, Africa
Shoebill (Balaeniceps rex) head portrait, Lake Victoria, Uganda, Africa
Shoebill (Balaeniceps rex) in swampland, head portrait, Mabamba, Lake Victoria, Uganda, Africa
Black triggerfish (Melichthys niger) swimming over reef, usually dark black in color, Hawaii, Pacific Ocean
Male Pygmy killer whale (Feresa attenuata) swimming in open ocean, Hawaii, Pacific Ocean
Spanish Imperial eagle (Aquila adalberti) perched on branch, looking down, Spain. February
Anadolu stallion standing alert, portrait, Sirince mountains, Turkey
Two Gentoo penguin (Pygoscelis papua) chicks sleeping huddled together, Port Lockroy, Goudier Island, Antarctic Peninsula, Antarctica
Tricolored heron (Egretta tricolor) fishing, taking off from water, . Everglades National Park, Florida, USA
Dorsal fin of Blacktip reef shark (Carcharhinus melanopterus) breaking water surface, at sunset, off island of Yap, Micronesia. Philippines sea, Pacific Ocean
Broad-bordered bee hawkmoth (Hemaris fuciformis) nectaring on Lavender (Lavandula sp. ) flower, Orvieto, Umbria, ItalyBroad-bordered bee hawkmoth (Hemaris fuciformis) nectaring on Lavender (Lavandula sp.) flower, Orvieto, Umbria, Italy. July
Black caiman (Melanosuchus niger) head half submerged in river, Yasuni National Park, Amazon Basin, Ecuador
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Puma (Puma concolor) walking with the Torres del Paine mountains in background, Torres del Paine National Park, Magallanes, Chile
Three King penguins (Aptenodytes patagonicus) reflections in water, Salisbury Plain, South Georgia, Southern Ocean
Two Canada geese (Branta canadensis) in flight with autumn foliage behind, Maryland, USA. October
Red grouse (Lagopus lagopus) standing on gritstone rock at sunrise, Peak District National Park, UK. August
Red grouse (Lagopus lagopus) stretching wings at sunrise, Peak District National Park, UK. August
Young Nile Crocodile (Crocodylus niloticus) basking on an exposed log, Chobe River, Botswana
Sylvia's tree frog (Cruziohyla sylviae), Veragua rainforest, Costa Rica
Male Australian magpie (Gymnorhina tibicen) standing on stump with beak full of moths, backlit. Canterbury, South Island, New Zealand
Hippopotamus (Hippopotamus amphibius) in shalow water with Cattle egret (Bubulcus ibis) in front of Mount Kilimanjaro, Amboseli National Park, Kenya
Puma (Puma concolor) female, walking in front of Torres del Paine massif, Torres del Paine National Park, Patagonia, Chile
Two Komodo dragons (Varanus komodoensis) males, fighting on a remote beach, Komodo Island, Komodo National Park, Indonesia. Endangered
Puma (Puma concolor) female, running in deep fresh, snow, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female with two cubs, aged six months, sitting in fresh snow, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) cub, aged nine months, walking in deep, fresh snow, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, walking in deep fresh, snow, Torres del Paine National Park, Patagonia, Chile
Common Darter (Sympetrum striolatum) in flight, Brasschaat, Belgium
Willow grouse (Lagopus lagopus) winter plumage, in flight, Utsjoki, Finland. February
King eider (Somateria spectabilis) male, on water, portrait, Batsfjord, Norway. February
Hen harrier (Circus cyaneus) juvenile male, in flight, Parainen Uto, Finland. September
Male Cheetah (Acinonyx jubatus) stretching and yawning on a fallen tree trunk, Okavango Delta, Botswana, Africa
American purple gallinule (Porphyrio martinica) leaping between water lily pads. Everglades National Park, Florida, USA
Western reef egret (Egretta gularis) wading in river at sunrise, Allahein River, The Gambia