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Red squirrel (Sciurus vulgaris) gnawing red deer antler for minerals, Cairngorms National Park, Highlands, Scotland, UK, June
Red squirrels (Sciurus vulgaris) interacting, Cairngorms National Park, Highlands, Scotland, UK, January
Scots pine (Pinus sylvestris) with Northern lights / Aurora borealis lighting up the night sky in background, Monadhliath Mountains, Cairngorms National Park, Scotland, UK, October 2015
Devils Marbles, Devils Marbles Conservation Reserve, Northern Territory, Australia, June 2010
Reflections in Llyn Gwynant on a misty morning, Glaslyn valley, with Yr Aran mountain in the background, Snowdonia National Park, North Wales, UK, November 2017
Yellow-plumed parakeets (Leptosittaca branickii) at nest cavity in wax palm stump, Tapichalaca Biological Reserve, Ecuador
Red Deer (Cervus elaphus) stag seen from behind a tree. Cairngorms National Park, Scotland, February
Bison (Bison bison) in snow. Yellowstone National Park, USA, February
Golden eagle (Aquila chrysaetos) adult female taking off, flying over mountain landscape, trained bird photographed during filming in Glenveagh National Park, Donegal, Ireland. August 2010
Broad-tailed Hummingbird {Selasphorus platycercus} male in flight feeding on Musk Thistle (Carduus nutans) Rocky Mountain National Park, Colorado, USA, June
Path through Heather flowering on lowland heathland, Rockford Common, Linwood, New Forest National Park, Hampshire, England, UK, dawn, August 2011
Great Staple Tor in evening light, Dartmoor National Park, Devon, UK
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Lava lake at night in the crater of Nyiragongo Volcano, Virunga National Park, North Kivu Province, Democratic Republic of Congo
Reflections in Llyn Gwynant in the Glaslyn valley, looking west with Yr Aran mountain in the background, Snowdonia National Park, North Wales, UK, June
RF- Carrion crow (Corvus corone) flying from a snow covered pine tree in a winter landscape. Valsavarenche, Gran Paradiso NP, Italy, March
Glanville fritillary butterfly (Melitaea cinxia) on blade of grass, Pollino National Park, Basilicata, Italy, May 2009
Creag Megaidh National Nature Reserve, Lochaber, Scotland, UK, December. 2020VISION Book Plate
The River Dee flowing past the Devils Point. Cairngorms National Park, Grampian Mountains, Scotland, UK, February 2012
Dartford warbler (Sylvia undata) perched on Heather twig, Arne RSPB reserve, Dorset, UK, July. 2020VISION Book Plate
Aerial view over Abernethy forest and mountains at dawn, Cairngorms National Park, Scotland, UK. April, 2020
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair playing with each other, one with hand over eyes of other, Manuel Antonio National Park, Quepos, Costa Rica
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Black rhino (Diceros bicornis) and White Rhino (Ceratotherium simum) bulls facing off. Solio Game Reserve, Laikipia, Kenya. September
Olive grove (Olea europaea) Vieste, Gargano National Park, Gargano Peninsula, Apulia, Italy, May 2008
Carboniferous limestone outcrops. Lathkill Dale National Nature Reserve, Peak District National Park, UK, June 2008
Mixed flock of Snow geese (Chen caerulescens atlanticus / Chen caerulescens) and Sandhill cranes (Grus canadensis) at wintering area at dawn, with full moon, Bosque del Apache, New Mexico, USA
Asiatic Wild Water Buffalo (Bubalus arnee), Royal Manas National Park, Bhutan
Female Eastern grey kangaroo (Macropus giganteus) standing next to joey, Grampians National Park, Victoria, Australia, May
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, lake Victoria, Uganda
Dry-stone walls and barns in Wensleydale, Yorkshire Dales National Park, North Yorkshire, England, UK, June 2016
Leaves of a Ginkgo tree or Maidenhair tree (Ginkgo biloba) Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Portrait of a meerkat (Suricata suricatta) or suricate, looking at the camera and standing to survey its territory on the edge of Makgadikgadi Pans National Park, Botswana, April
RF- European grey wolf (Canis lupus) running through snow in birch forest, Tromso, Norway. Captive, April
Sri Lankan elephant (Elephas maximus maximus) walking, Yala National Park, Southern Province, Sri Lanka
Red tailed boa constrictor (Boa constrictor) on fallen tree over water, Yasuni National Park, Orellana, Ecuador
Eastern green mamba {Dendroaspis angusticeps} Arabuko-Sokoke forest, Kenya
Young Ring-tailed lemur (Lemur catta) 6-8 weeks, clinging to mother, Berenty Private Reserve, southern Madagascar, November
Fafiao Waterfalls, Peneda Geres National Park, Portugal, February 2009
Red fox (Vulpes vulpes) silhouetted in mixed beech-Douglas fir forest. Freiburg, Black Forest National Park, Germany, October
English oak tree (Quercus robur) standing in a formerly farmed meadow with many flowering Meadow buttercups (Ranunculus acris) surrounded by mature hedgerows
Yellowstone River. Yellowstone National Park, Wyoming, USA. September
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Granitic boulders at Karlu Karlu / Devils Marbles Conservation Reserve, framed by Eucalyptus tree trunks. Northern Territory, Australia. 2008
Sun rays shining through clouds onto Loughrigg Fell and hills of the Lake District National Park, in evening. Ambleside, Cumbria, England, UK. April 2020