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Dobsonfly female (Corydalinae) in cloud forest, Manu Biosphere Reserve, Peru
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
White witch moth (Thysania agrippina) camouflaged against bark of tree. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
RF - Maidens veil / Bridal veil fungus (Phallus indusiatus) with indusium fully formed, on rainforest floor. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Tufted / Brown capuchin (Cebus apella), female with baby on back, sitting in tree, mid-altitude montane forest, Manu Biosphere Reserve, Peru
Grey whale (Eschrichtius robustus) breaching, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Wild boar (Sus scrofa) eyes glowing at dusk, introduced species in La Pampa, Argentina
Warthogs (Phacochoerus aethiopicus) mother with babies drinking, Mkhuze game reserve, KwaZuiu-Natal, South Africa, June
Pygmy Hog (Porcula salvania) foraging. Captive, member of a reintroduction project. Assam, eastern India
Grykes in limestone pavement at sunset, , Twisteleton Scar End, Yorkshire Dales National Park, Yorkshire, England, UK, May 2013
Bubble Tip Anemone (Entacmaea quadricolor). The tentacles are usually a dull brown, but this anemone is partially bleached and has lost most of the endosymbiotic zooxanthellae
Leather Coral (Sinularia flexibilis). Komodo National Park, Indonesia
European roller (Coracias garrulus) peering out of nest with feather, Lake Csaj, Pusztaszer, Kiskunsagi National Park, Hungary, February
African wild dog (Lycaon pictus) portrait, Mana Pools National Park, Zimbabwe
Elephant (Loxodonta africana), male standing on hind legs to reach acacia pods with Cattle egrets (Bubulcus ibis) in background. Mana Pools National Park, Zimbabwe
Bee-eater (Merops apiaster) with Bumblebee prey, Sierra de Grazalema Natural Park, southern Spain, July
Crab spider (Thomisus onustus) Sierra de Grazalema Natural Park, southern Spain. May
Iberian lynx (Lynx pardinus) with Magpie (Pica pica) behind, Sierra de Andujar Natural Park, Jaen, Spain, September
Lava lizard (Microlophus peruvianus) juvenile peering out of eye of dead sea lion. Paracas National Reserve, Peru. Highly commended in the Animals in their Environment Category of Wildlife
Mountain hare (Lepus timidus) in extreme winds with spin drifts of snow, Cairngorms National Park, Scotland, UK, February 2018
Blue butterfly (Lycaenidae) frozen in ice, Cortes de la Frontera, Los Alcornocales Natural Park. Spain
Marine otter (Lontra felina) female and cubs on coastal rocks, Paracas National Reserve, Peru, Endangered species
Marine otter (Lontra felina) female on coastal rocks, Paracas National Reserve, Peru, Endangered species
Scarletina bolete (Boletus luridiformis) toadstool, New Forest National Park, Hampshire, England, UK. September
Mozambique Spitting Cobra (Naja mossambica) ejecting venom Kruger, South Africa, Controlled conditions
Ammonite fossil on beach, Charmouth, Jurassic Coast World Heritage Site, Dorset, England, November 2004
Tibetan macaque (Macaca thibetana) Tangjiahe Nature Reserve, Sichuan Province, China
Springbok (Antidorcas marsupialis) herd fleeing predator, Kgalagadi Transfrontier Park, South Africa
RF - Meerkat (Suricata suricatta) resting on cool sand, Kgalagadi Transfrontier Park, South Africa
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Leaves of a Ginkgo tree or Maidenhair tree (Ginkgo biloba) Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Red-billed blue magpie (Urocissa erythrorhyncha) perched on branch, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Amazon kingfisher (Chloroceryle amazona) with fish, Cuiaba, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male, hunting Capybara (Hydrochoerus hydrochaeris). The capybara while jumping away has cut the nose of the Jaguar with its toemail
Jaguar (Panthera onca) male, chasing a Caiman. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Wild female Amur leopard (Panthera pardus orientalis) looking up, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
Lioness (Panthera leo) resting up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) cub up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) male standing in cold and rain with strong wind blowing, smelling the air, Masai Mara National Reserve, Kenya
African lion (Panthera leo) young male stalking towards camera, Etosha National Park, Namibia October
African lion (Panthera leo) pair, Masai Mara National Reserve, Kenya, January
Male Lion running through grass {Panthera leo} Masai Mara, Kenya
Female Bornean orangutan (Pongo pygmaeus) sitting in a tree, Tanjung Puting National Park, Borneo-Kalimantan, Indonesia
