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Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
Atlantic Puffin (Fratercula artica) adult on breeding cliffs. Hermaness NNR, Shetland, UK, June
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Young Polar bear (Ursus maritimus) standing and trying to balance in shallow water along the Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska
Plains zebra (Equus quagga) grooming foal, Kruger National Park, South Africa, July
Snow leopard (Panthera uncia) female in snow, Hemis National Park, Ladakh, India
Brown bear (Ursus arctos) male, portrait. Martinselkonen, Kainuu, Finland. June
Bamboo lemur (Hapalmur griseus) in tree. Andasibe-Mantadia National Park, Madagascar
Short-beaked echidna (Tachyglossus aculeatus). Francois Peron National Park, Shark Bay, Western Australia. October
Painted mantella frog (Mantella madagascariensis) Mantadia NP, Eastern Madagascar
Blue poppy (Meconopsis zangnanensis) Mt Qomolangma National Park, Qinghai Tibet Plateau, China
Arctic fox (Vulpes lagopus) portrait of juvenile, winter pelage. Dovrefjell National Park, Norway. September
Arctic fox (Vulpes lagopus), juvenile running through snow, winter pelage. Dovrefjell National Park, Norway. September
Bearded pig (Sus barbatus) close up of snout, Bako National Park, Sarawak, Borneo
European swallowtail butterfly (Papilio machaon gorganus) on flower, Mercantour National Park, Provence, France, June
Vale of Ewyas, view over Capel-y-ffin and the valley of the Afon Honddu, Black Mountains, Brecon Beacons National Park, Powys, Wales, UK, October 2008
Mountain gorilla (Gorilla gorilla beringei) silverback Gihishamwotsi displaying, non group dominant, Sabyinyo Group, Volcanoes National Park, Virunga Mountains, Rwanda
Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Rainbow river, Rainbow Springs State Park, Florida, USA. January. Photographed for The Freshwater Project
Landscape of Zanskar River, a tributary of the Indus, borders part of Hemis national park, habitat of the snow leopard (Panthera uncia), Hemis National Park, Ladakh, India
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Bison (Bison bison) in snowstorm. Yellowstone National Park, USA, February
Glanville fritillary butterfly (Melitaea cinxia) on blade of grass, Pollino National Park, Basilicata, Italy, May 2009
Field of Meadow saffron crocus (Colchicum autumnale) one growing in cow dung, Mohacs, Bda-Karapancsa, Duna Drava NP, Hungary, September 2008
Muskox (Ovibos moschatus) with snow on face, Dovrefjell National Park, Norway, February 2009. WWE INDOOR EXHIBITION
Arctic tern (Sterna paradisaea) in flight, Inner Farne, Farne Islands, Northumberland, June
Mallard (Anas platyrhynchos) male flying low over water. Cambridgeshire Fens, England, March
Caledonian pine forest in mist at sunrise, Rothiemurchus Forest, Cairngorms NP, Highland, Scotland, UK, June 2011. 2020VISION Exhibition. 2020VISION Book Plate
New Forest pony grazing on Latchmore Bottom at dawn, view from Dorridge Hill, The New Forest National Park, Hampshire, England, UK, July 2011. 2020VISION Book Plate
Glen Affric in autumn, Highlands, Scotland, UK, October 2012
Looking over Rothiemurchus ancient Caledonian pine forest at dawn. Cairngorms National Park, Highlands, Scotland, UK, September 2015
Sheep sheltering from harsh weather behind a stone wall, Peak District National Park, Derbyshire, UK. March
Grass Snake (Natrix natrix) portrait with tongue extended, Staffordshire, England, UK, April
Brown throated Three-toed Sloth (Bradypus variegatus) mother and newborn baby (less than 1 week old) Aviarios Sloth Sanctuary, Costa Rica *Digitally removed highlights in background
Alpine marmot (Marmota marmota), with Mount Grossglockner (3798m) in background, Hohe Tauern National Park, Austria, July
Silver Studded Blue (Plebejus argus), Gran Paradiso National Park, Aosta Valley, Pennine Alps, Italy. July
Ogre faced / Net-casting spider {Deinopis sp} with web held between legs that it will stretch over prey that walks below it. Masoala Peninsula National Park, north east Madagascar
Bornean horned frog (Megophrys nasuta) on forest floor. Danum Valley, Sabah, Borneo, Malaysia
Pied avocet (Recurvirostra avosetta) feeding in shallow water at dawn, Etosha National Park, Namibia
handsAlpine marmot (Marmota marmota) reching upwards, Hohe Tauern National Park, Austria, July
Male Saiga antelope (Saiga tatarica) in winter, The Black Lands National Park, Kalmykia, Russia
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair close-up Manuel Antonio National Park, Quepos, Costa Rica
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Male silverback Eastern lowland gorilla (Gorilla beringei graueri) in the equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Silverback male Eastern lowland gorilla (Gorilla beringei graueri) intimidation display by chest beating, equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka climbing a tree, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
