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Ancient Scots pine (Pinus sylvestris) amongst flowering Common heather / Ling (Calluna vulgaris). Rothiemurchus Forest, Cairngorms National Park, Scotland, UK
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Eurasian badger (Meles meles), two foraging in Pine (Pinus sp) woodland. Glenfeshie, Cairngorms National Park, Scotland, UK
Rocks and coastal rainforest. Cocos Island National Park, Costa Rica. 2018
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Fish seeking shelter around floating tree in open ocean. Off coast of Cocos Island National Park, Costa Rica
Central American spider monkey (Ateles geoffroyi) juveniles hanging by tails and playing, Calakmul Biosphere Reserve, Yucatan Peninsula, Mexico, August
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Cuban crocodile (Crocodylus rhombifer) in a cenote in Cienaga de Zapata National Park. Cuba. Critically endangered species
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
One-horned Asian rhinoceros (Rhinoceros unicornis), Chitwan National Park, Inner Terai lowlands, Nepal
Arctic fox (Vulpes lagopus), juvenile licking lips, winter pelage. Dovrefjell National Park, Norway. September
Indian rhinoceros (Rhinoceros unicornis) Kaziranga National Park, Assam, India
Melanistic leopard / Black panther (Panthera pardus) Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Young orphan Elephants (Loxodonta africana) kissing. David Sheldrick Wildlife Trust Nairobi Elephant Nursery, Kenya, July 2010
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Red fox (Vulpes vulpes) resting with eyes open, Vanoise National Park, Rhone Alpes, France. October
Flowers (Trollius sp) Balang Mountain, Wolong National Nature Reserve, Sichuan Giant Panda Sanctuaries - Wolong, Mt Siguniang and Jiajin Mountains UNESCO World Heritage Site, Sichuan Province, China
Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Low angle view of Porcelain fungus (Oudemansiella mucida) growing on a dead Beech tree (Fagus sylvatica). Plitvice Lakes National Park, Croatia. November
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June
Giant sequoia (Sequoiadendron giganteum) trees in Sequoia National Park, California, USA, September 2014
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Close up if a Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Young Polar bear (Ursus maritimus) standing on hing legs, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Cave Pearls, formed as dripping water rich in calcium salts deposits calcite around a small nucleus (often a sand grain). The constant movement of each cave pearl by the dripping water keeps them
Bacteria - iron and manganese oxidizers and their complementary reducers, in the mud of an empty fish pond, Upper Lusatia Heath and Pond Landscape, Upper Lusatia UNESCO Biosphere Reserve, Germany
Rowan (Corbus aucuparia) and Ash trees (Fraxinus excelsior) in fruit, blurred by zoom effect, Glenfeshie, Cairngorms National Park, Scotland, UK, October
Aerial view river tributaries and saltmarshes of Bahia de Cadiz Natural Park, Huelva, Andalucia, Spain, March 2008
Detail of Ocellated Turkey (Meleagris / Agriocharis ocellata) plumage. El Mirador-Rio Azul National Park, Department of Peten, Guatemala
Small tortoiseshell butterfly (Aglais urticae) feeding on Ice plant (Sedum sp. )Small tortoiseshell butterfly (Aglais urticae) feeding on Ice plant (Sedum sp.). Peak District National Park, Derbyshire, UK. September
Yunnan, or Black Snub-nosed monkeys (Rhinopithecus bieti) adult and young, Ta Cheng Nature reserve, Yunnan, China, October
Red beetle (Pyrochroidae) sitting on a leaf, Tongbiguan Nature Reserve, Dehong prefecture, Yunnan province, China, May
Yunnan Snub-nosed monkey, (Rhinopithecus bieti) portrait, Ta Chen NP, Yunnan province, China
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
River running through cloud forest, Manu Biosphere Reserve, Amazonia, Peru
Dobsonfly female (Corydalinae) in cloud forest, Manu Biosphere Reserve, Peru
Amazonian Horned Frog (Ceratophrys cornuta) camouflaged amongst leaf litter on lowland rainforest floor, waiting to ambush passing prey. Manu Biosphere Reserve, Amazonia, Peru. November
Giant waxy monkey frog (Phyllomedusa bicolor). Lowland Amazon rainforest, Manu Biosphere Reserve, Peru
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
White witch moth (Thysania agrippina) camouflaged against bark of tree. Manu Biosphere Reserve, Amazonia, Peru