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Lenticular clouds above Torres del Paine National Park. Patagonia, Chile. November 2018
Guanaco (Lama guanicoe) female, alert whilst looking at Puma (Puma concolor) on on opposite slope, portrait. Torres del Paine National Park, Patagonia, Chile. November
Giant river otter (Pteronura brasiliensis) resting on fallen tree trunk above Cuiaba River. Pantanal, Mato Grosso, Brazil
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Spanish lavender (Lavandula stoechas). Faia Brava Reserve, Archaeological Park of the Coa Valley, Western Iberia, Portugal. April
Melanistic leopard / Black panther (Panthera pardus fusca) resting in tree. Nagarhole National Park, India. Photo© Phillip Ross/Felis Images
Asiatic lion (Panthera leo persica), two females and two cubs. Gir National Park, Gujarat, India. Photo© Phillip Ross/Felis Images
Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia
Western lowland gorilla (Gorilla gorilla gorilla) male aged 12 years standing up at edge of Mpassa river. Captive bred in UK and released into wild along with his father in 2013
Sacred lotus (Nelumbo nucifera) flowering in Puzhehai Lake with peaks in background. Puzhehei National Wetland Park, Yunnan Province, China. 2009
Laurisilva Forest, trees covered in lichen / moss at Terceira Island, Azores, August 2011
Aardvark (Orycteropus afer) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa. Medium repro only
Tropical shield mantis (Choeradodis sp. ) Sumaco National Park, Napo, EcuadorTropical shield mantis (Choeradodis sp.) Sumaco National Park, Napo, Ecuador
Giant river otter (Pteronura brasiliensis) eating a catfish. Yasuni National Park, Orellana, Ecuador
Spangle waxcap (Hygrocybe insipida) Peatlands Park, County Armagh, Northern Ireland, October
Red Fox (Vulpes vulpes) portrait in autumn grass, with Kronotsky Volcano on the horizon. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, August
Robins pincushion gall caused by Gall wasp (Diplolepis rosae) on wild Dog rose (Rosa canina), Peak District National Park, Derbyshire, UK. September
Small lochan surrounded by flowering heather (Ericaceae sp) on the edge of the Caledonian pine forest, Rothiemurchus, Cairngorms NP, Scotland, UK, September 2011
Bengal tiger (Panthera tigris tigirs) tigress, stopping due to sound from camera while patrolling, Kanha National Park, Central India. Camera trap image
Bengal tiger (Panthera tigris tigris) (T27) walking on bund / dam wall trail. Kanha National Park, Central India. Camera trap image
Tiger (Panthera tigris tigris) dominant male tiger approaching Beja pani waterhole, Kanha National Park, Central India. Camera trap image
Indian Flying Fox (Pteropus giganticus) male roosting in tree, Keoladeo NP, Bharatpur, India
Rainbow lorikeet (Trichoglossus moluccanus) on a branch, Cania Gorge National Park, Queensland, Australia. September
Mountain hare (Lepus timidus) in summer pelage amongst heather, Cairngorms National Park, Scotland, UK, August
Curled octopus (Eledone cirrhosa) swimming over sea floor, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
Peacock butterfly (Inachis io) feeding on Field Scabious (Knautia arvensis) in a disused limestone quarry. Peak District National Park, Derbyshire, UK. August
Tufted duck (Aythya fuligula) male drake, Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July
Great crested grebe (Podiceps cristatus) adult with young on its back, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June
Mute swan (cygnus olor) family, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Alpine ibex (Capra ibex) struggling in deep snow on a steep slope, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, January
Tayra (Eira barbara) climbing in a tree in rainforest habitat with fern covered trees, Bellavista private reserve, Mindo cloud forest area, Ecuador, July
Mountain gorilla (Gorilla beringei) juvenile aged 2 years, hanging from branch, member of the Nyakagezi group, Mgahinga National Park, Uganda. January
Phuket horned tree lizard (Acanthosaura phuketensis) near Bang Pae Waterfall, Khao Phra Thaew NP, Phuket Island, Thailand
Autumnal beech trees ((Fagus sylvatica) at Bolderwood, The New Forest, Hampshire, UK. November
Clapper Bridge at Postbridge, early morning light and mist, Dartmoor National Park, Devon, UK. October 2018
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
European otter (Lutra lutra) two at waters edge, one shaking off water, Kiskunsagi National Park, Pusztaszer, Hungary, October
European otters (Lutra lutra) feeding at water?s edge with splashing water, Kiskunsagi National Park, Hungary, January
Lone Scots pine tree (Pinus sylvestris) and star trails with the north star, Abernethy forest, Cairngorms National Park, Scotland, UK
Asian black bear (Ursus thibetanus) feeding on leaves, Tangjiahe Nature Reserve, Sichuan Province, China, April
Tibetan wolf (Canis lupus chanco) in mountain landscape, Serxu County, Garze Prefecture, Sichuan Province, China
Snow leopard (Uncia uncia) with Bharal (Pseudois nayaur) prey, Serxu County, Garze Prefecture, Sichuan Province, China
Tibetan macaque (Macaca thibetana) carrying young baby, Tangjiahe Nature Reserve, Sichuan Province, China
Sumatran Orangutan (Pongo abelii) female with very young baby, Gunung Leuser National Park, Sumatra, Indonesia
Orangutan {Pongo pygmaeus} adult climbing vine, Rehabilitation sanctuary, Tanjung Puting National Park, Kalimantan, Indonesia