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Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Veragua cross-banded treefrogs (Smilisca sordida) mating pair at night, Osa Peninsula, Costa Rica
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Young Sperm whale (Physeter macrocephalus) with erect penis Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Long-tailed manakin (Chiroxiphia linearis) perched on lek (display perch), Costa Rica
Orange tip butterfly (Anthocharis cardamines) female and male, visiting Honesty flower (Lunaria annua) Wiltshire, England, UK, May
Sooty Albatross (Phoebetria fusca) group courting on high inland ridge. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endangered
RF - Red deer (Cervus elaphus) stags fighting, Holkham Park, Norfolk, England, UK, September
Black Grouse (Tetrao tetrix) lek with male displaying and females around in winter, Tver, Russia. April
Black Grouse (Tetrao tetrix) females in tree in winter, Tver, Russia. April
Black Grouse (Tetrao tetrix) males fighting in winter, Tver, Russia. April
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Female Black veined white butterfly (Aporia crataegi) laying eggs on Buckthorn, Lagadin region, Lake Ohrid, Galicica National Park, Macedonia, June 2009
Nudibranch eggs, Larvotto Marine Reserve, Monaco, Mediterranean Sea, July 2009
Nudibranch (Adalaria proxima) egg laying, Moere coastline, Norway, February 2009
Female Loggerhead turtle (Caretta caretta) crawling up beach in search of spot to lay eggs, Dalyan Delta, Turkey, August 2009
Black grouse (Tetrao tetrix) male displaying at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Brown hare (Lepus europaeus) two males chasing female in snow covered field during a blizzard. Derbyshire, England, UK, February. Winner of British Seasons category of BWPA competition 2014
Moorland Clouded Yellow butterfly (Colias palaeno) male and female, Joutsa (formerly Leivonmaki), Finland, July
Silver-studded blue butterfly (Plebejus argus) pair mating, resting on a Pyrimidal orchid (Anacamptis pyramidalis) flower, Gwithian Towans, Cornwall, UK, July
Green Turtle (Chelonia mydas) female on beach, Concha Beach, municipality of Itacare, Southern Bahia State, Eastern Brazil
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Gannet nesting colony on Bass Rock, UK
Indigo hamlets (Hypoplectrus indigo) pair spawning at dusk, George Town, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. Shark City, Ulong, Rock Islands, Palau
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Atlantic salmon (Salmo salar) leaping upstream in river, Scotland, UK