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Eared grebes (Podiceps nigricollis) stand erect in water during courtship, North Park, Colorado, USA. June
Coscoroba swan (Coscoroba coscoroba) at nest with eggs. La Pampa, Patagonia, Argentina. February
Seep monkey flower (Mimulus guttatus), bifid stigma above stamens. Nectar spot guides and central groove leading pollinators to nectar. Focus stacked
Snowdrop (Galanthus nivalis) anthers and inner petals. Outer petals have been removedSnowdrop (Galanthus nivalis) anthers and inner petals.Outer petals have been removed. Focus stacked
Himalayan balsam (Impatiens glandulifera), dissection of flowers. Male phase above, downward projecting stamens release pollen onto backs of bumblebees and honey bees
Hairy banana (Musa velutina), self peeling exposing flesh to aid seed dispersal by birds. In glasshouse. Surrey, England, UK
Scarab beetle (Scarabaeoidea) pair mating inside Tulip (Tulipa micheliana) flower. Beetles covered in pollen. Uzbekistan, April
Bumblebee scarab (Amphicoma vulpes) pair mating in Tulip (Tulipa armena) flower. Pollen picked up on beetles and transferred to other flowers. Near Karabel Pass, Turkey. May
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
Mauritian bloody bellflower (Nesocodon mauritianus), female phase with red nectar at base of style. Cultivated at Kew Gardens, London, England, UK. Native to Mauritius
Fallow deer (Dama dama) buck bellowing in autumn vegetation, Bradgate Park, Leicestershire, UK, October
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Sparkling violetear hummingbird (Colibri coruscans) incubating eggs at nest, Andean cloud forest, Ecuador
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Chum salmon (Oncorhynchus keta) migrate up a small river near Bella Bella, British Columbia, Canada. September
Seven spot ladybird (Coccinella septempunctata) pair mating on Mayweed (Anthemis sp) flower. Cyprus. April
Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Western Grebe (Aechmorphus occidentalis), pair during rushing courtship display in which they run across waters surface in synchrony, near Escondido, California, USA, January
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Jaguars (Panthera onca) courting pair on a sand bank. Cuiaba River, Northern Pantanal, Mato Grosso, Brazil
Two rainbow lorikeets (Trichoglossus moluccanus) on branch, Werribee, Victoria, Australia
Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Clarion Lizard (Urosaurus clarionensis) displaying, IUCN Vulnerable, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname
Bignose unicornfish (Naso vlamingii) profile of male displaying (the blue stripe at the bottom of the frame is the tail of another individual) Baa Atoll, Maldives. Indian Ocean
Hoopoe (Upupa epops) male feeding female in courtship display, Pusztaszer, Hungary, April
Iolas blue butterfly (Iolana iolas) pair mating, Aosta Valley, Gran Paradiso National Park, Italy
Iolas blue butterfly (Iolana iolas) pair mating on flowers, Aosta Valley, Gran Paradiso National Park, Italy
European bee eater (Merops apiaster) pair at nest at breeding site, Danube delta rewilding area, Romania, June
Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Nudibranchs mating (Polycera faeroensis) Sark, British Channel Islands, July
Red deer (Cervus elaphus) stag and two hinds, Los Alcornocales Natural Park, southern Spain, September
European bee eaters (Merops apiaster) mating, Sierra de Grazalema Natural Park, El Bosque, southern Spain, May
Underwater view of Toads (Bufo bufo) and toadspawn in lake, Ain, Alps, France
Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India
Kottigehara dancing frog (Micrixalus kottigeharensis), dancing frog name given because of foot flagging behavior between competing males. Endemic to Western Ghats. Agumbe, Karnataka, India
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Whitebelly damselfish (Amblyglyphidodon leucogaster) pair spawning / laying eggs on a seafan (Annella mollis). Ras Mohammed National Park, Sinai, Egypt. Red Sea
Skeleton shrimp (Caprella septentrionalis) mating Novaya Zemlya, Russian Arctic, July
