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White-lipped deer (Cervus albirostris) walking on the Tibetan plateau, Serxu, Garze Prefecture, Sichuan Province, China
Clearfin lionfish (Pterois radiata) mating. The female (left) has just released a raft of eggs, which is visible between her tail and the red face of the male
Portrait of a Blackbar soldierfish (Myripristis jacobus) infected with Cymothoid isopod parasites. The large parasite between the eyes is a female, the smaller individuals are males
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Azure / common coenagrion damselflies (Coenagrion puella) pair mating, Meeth Quarry, Devon, UK. July
Sockeye salmon (Oncorhynchus nerka) migration, Adams River, British Columbia, Canada. October
Green sea turtle (Chelonia mydas) heading back to sea after laying eggs on the beach, sunrise, Ras Al-Jinz Turtle Reserve, Ash-Sharkiyah, Oman, November
Galapagos hawk (Buteo galapagoensis) landing on mating pair of Galapagos giant tortoise (Chelonoidis vandenburgh) Alcedo Volcano, Isabela Island, Galapagos
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Common Merganser (Mergus merganser) female with ducklings, in threat posture at the approach of a second female, Lansing, New York, USA
Leopard (Panthera pardus) pair mating behaviour, Okavango Delta, Botswana
Black Grouse (Lyrurus tetrix) fighting on icem Finland. March
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Eurasian rollers (Coracias garrulus) courting on branch, Hungary. June
Small emerald spreadwing (Lestes virens) pair mating. Klein Schietveld, Brasschaat, Belgium. August
Blue-tailed damselfly (Ischnura elegans) pair mating on stem. Brasschaat, Belgium. August
King cormorants (Phalacrocorax atriceps) in courtship behaviour at their nest site. Saunders Island, Falkland Islands
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Crocus (Crocus sp) flower from above, closing in evening. Focus stacked
Water tulip (Cypella aquatilis) from above, in garden pond. Native to Brazil
Peach leaved bellflower (Campanula persicifolia) dissection, petals removed to show how stamens transfer pollen to style via secondary pollen presentation in bud stage. Focus stacked
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Common cranes displaying by woodland {Grus grus} Finland
Elephant yam (Amorphophallus paeoniifolius) in UV light, lower band with female flowers, male flowers above
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Fallow deer (Dama dama) herd at dawn, Holkham, Norfolk, England, UK, October
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Red deer (Cervus elaphus) herd of female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) female does and young crossing water, Jura, Scotland, UK, September
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Magellanic penguin (Spheniscus magellanicus) calling at the entrance of burrow, courtship display, Carcass Island, Falkland Islands, October
Green Turtle (Chelonia mydas) returning to the sea, IUCN Endangered, Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Common toad (Bufo bufo) pair in amplexus, Deeside, Scotland April
Iolas blue butterfly (Iolana iolas) pair mating on flowers, Aosta Valley, Gran Paradiso National Park, Italy
Brown Hare (Lepus europaeus), Bayern, Germany. April
Red snappers / Two-spot red snappers (Lutjanus bohar) mass shoal of fish spawning, Palau, Philippine Sea. A few black and white snappers (Macolor niger) have crawled into the school to eat the eggs
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Veragua cross-banded treefrogs (Smilisca sordida) mating pair at night, Osa Peninsula, Costa Rica
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
Young Sperm whale (Physeter macrocephalus) with erect penis Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
