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Skeleton shrimp (Caprella septentrionalis) mating Novaya Zemlya, Russian Arctic, July
Western capercaillie (Tetrao urogallus) male calling at lek, with female, Tver, Russia. May
Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April
Male Red deer bellowing during rut. Denmark, Jaegersborg Dyrehaven Copenhagen
Gunnison sage-grouse (Centrocercus minimus) male displaying at a lek. Gunnison County, Colorado. April
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Parasitoid wasp (Ichneumonidae) carrying a large paralysed caterpillar to its nesting hole, The Peloponnese, Greece, May 2009
European grayling (Thymallus thymallus) pair spawning, the dark one is the male, Lake Thun, Thun, Switzerland, March 2009
Whooper swan (Cygnus cygnus) pair on loch in heart shape, Cairngorms National Park, Scotland, UK, November
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
Common dolphins (Delphinus delphis) pair in mating ritual, Atlantic ocean, Portugal, July
Woodcock orchid (Ophrys cornuta / scolopax) with mating Black veined white butterfliesWoodcock orchid (Ophrys cornuta/scolopax) with mating Black veined white butterflies (Aporia crataegi), Bela Reka, Eastern Rhodope Mountains, Bulgaria, May 2013
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Tree frog (Ostocephalus oophagus) in bromeliad, female lays eggs for her tadpoles to eat. Brazil, South Americal
A Mountainous star coral (Montastraea faveolata) spawning at night, with gamete bundle being released from a coral polyp, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Atlantic salmon (Salmo salar) leaping upstream in river, Scotland, UK
Olive ridley turtle emerging from sea at dusk. Costa Rica {Lepidochelys olivacea}
Yellow-eyed penguin in forest {Megadyptes antipodes} Enderby Island, Auckland Is, New Zealand
Shelled Pteropod / Sea butterfly {Diacria trispinosa} with egg string attached, from between 188 and 507m, Mid-Atlantic Ridge, North Atlantic Ocean
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Three Common toads (Bufo bufo) in amplexus, Lucerne, Switzerland. March. Cropped
Common toad (Bufo bufo) and Common frog (Rana temporaria) in attempted amplexus, Lucerne, Switzerland. April
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October
Pair of Waved albatrosses (Phoebastria irrorata) billing as greeting. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Wine throated hummingbird (Selasphorus ellioti) male, courtship display, Cloud forest, Atitlan Volcano, Solola, Guatemala
Ruddy duck (Oxyura jamaicensis) male in courtship display. Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Leatherback turtle (Dermochelys coriacea) female, returning to sea after laying eggs in nest on beach, Grande Riviere, Trinidad Island, Trinidad & Tobago, Caribbean
Humpback whales (Megaptera novaeangliae) pair engaging in courtship behavior, dancing for hours, off Tubuai, French Polynesia, Pacific Ocean
Bluethroat (Luscinia svecica cyanecula) male, singing as a courtship display, Germany
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Magnificent frigatebirds (Fregata magnificens) pair in courtship display, male with gular sac inflated, North Seymour Island, Galapagos National Park, Galapagos Islands
Club-winged manakin (Machaeropterus deliciosus) male, courtship display at lek, Choco rainforest, Pichincha, Ecuador
Pair of Broadclub cuttlefish (Sepia latimanus) mating in a coral reef, Kagoshima Prefecture, Japan. Having succeeded in gaining the female's acceptance
Spawning Leaf corals (Pavona decussata). Spawning is synchronized over a large area with many coral colonies, Kyushu, Japan, Pacific Ocean. July
Female Northern pygmy squid (Idiosepius paradoxus) preparing to lay another egg on a blade of eelgrass (Zostera marina), Yamaguchi Prefecture, Japan
Male Korean seahorse (Hippocampus haema) giving birth. Kumamoto Prefecture, Kyushu, Japan
Fosa (Cryptoprocta ferox) pair, mating in forest canopy, Zombitse-Vohibasia National Park, central south western Madagascar
European Common adder (Vipera berus) pair courting and retreating under gorse (Ulex sp). Buckinghamshire, England, UK. May
Pair of Red mason bees (Osmia bicornis) mating on wallflower in garden. Hertfordshire, England, UK. May. Focus Stacked
Brown hare, (Lepus europaeus), male and female displaying courting behaviour, Islay, Scotland, UK. March
Brown hare, (Lepus europaeus), males in pursuit of female that is in season, Islay, Scotland, UK. March
Pair of Spotted unicornfish (Naso brevirostris) courtship display above the reef, Ari Atoll, Maldives, Indian Ocean
