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Brimstone butterfly (Gonepteryx rhamni) resting among foliage, Meeth Quarry, Devon, UK. June
Marbled White butterfly (Melanargia galathea) resting on flower head, Dunsdon Nature Reserve, Devon, UK. July
Male common blue butterfly (Polyommatus icarus) basking wings open on grass, Vealand Farm, Devon, UK. June
Gila woodpecker (Melanerpes uropygialis), emerging from nest in Saguaro cactus, Arizona, USA. July
Portrait of a female rescued Grey-headed flying-fox (Pteropus poliocephalus) pup. Captive - Taken in controlled conditions and under supervision of wildlife carer
Heath fritillary butterfly (Melitaea athalia) on daisy flower, Pyrenees National Park, France, June
Baikal seal (Pusa sibirica) underwater, endemic to Lake Baikal, Siberia, Russia. July
Feral domestic cattle, Puerto Fragata, Isabela Island, Galapagos
A Bottlenose dolphin (Tursiops truncatus), heavily scarred, possibly from shark attacks, on sand bank off Eleuthera, Bahamas
Alder buckthorn (Frangula alnus) in grassland at sunrise. Klein Schietveld, Brasschaat, Belgium. August 2019
Common Cuckoo (Cuculus canorus) perched on Cherry tree blossom Surrey, England, UK. April
Super Blood Wolf Moon or Blood Moon Eclipse so called because a Lunar eclipse is appearing at the same time as a Supermoon. Arundel, West Sussex, England, UK, 21 January 2019
Manchurian crane (Grus japonensis) pair in courtship dance during snowstorm. Hokkaido, Japan. March
Ezo red fox (Vulpes vulpes schrencki) sitting on snow, licking lips after eating Vole. Hokkaido, Japan. April
Humpback whale (Megaptera novaeangliae australis) female, stationary with calf heading to surface for breath of air. Vava'u, Tonga, South Pacific
Southern humpback whale (Megaptera novaeangliae australis), fluke of male as he dives. Vava'u, Tonga, South Pacific
Atlantic puffin (Fratercula arctica) swimming on the ocean surface, Svalbard, Norway, June
Polar bear (Ursus maritimus) female with a single young cub, only a few months old, northern Svalbard, Norway, June
Cuttlefish (Sepia officinalis) feeding on Scorpionfish (Scorpaenidae). Tenerife, Canary Islands
Blue shark (Prionace glauca). Canary Islands. North Atlantic Ocean
Arctic fox (Alopex lagopus) following scent, in winter pelage. Svalbard, Norway. April
Arctic fox (Alopex lagopus) standing in tundra. Svalbard, Norway, July
Mute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April
Mute swan (Cygnus olor) swimming with cygnets in misty lake in morning light. Richmond Park, London, England, UK. May. Cropped image
Mute swan (Cygnus olor) cygnet swimming, cocking head. Barnes, London, England, UK. April
California condor (Gymnogyps californianus) in flight, wings radio tagged. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. 2011
Indian leopard (Panthera pardus fusca) on rock with forested hills beyond. Nilgiri Biosphere Reserve, India. Camera trap image. 2019
Honey bee (Apis mellifera) extending proboscis to nectar on Common rue (Ruta graveolens). Green aphid (Aphidoidea) beneath flower. Surrey, England, UK. July
Snowdrop (Galanthus nivalis) anthers and inner petals. Outer petals have been removedSnowdrop (Galanthus nivalis) anthers and inner petals.Outer petals have been removed. Focus stacked
Snowdrop (Galanthus nivalis) dispersing pollen by sonication which replicates buzz pollination. Controlled conditions
Chilean bellflower (Crinodendron hookerianum) releasing pollen via sonication which replicates buzz pollination. Native to Chile. Controlled conditions
New Zealand tree fuchsia (Fuchsia excorticata) with blue pollen and yellow tipped stigma. Focus stacked
Bloody bellflower (Nesocodon mauritianus), female phase with red nectar surrounding stigma. Native to Mauritius. Cultivated at Kew Gardens, London, England, UK
Morning glory (Ipomoea tricolor) flower with pollen grains scattered by visiting insects
Celandine crocus (Crocus korolkowii) carpet in area of snowmelt. Pollinated by solitary bees and bumblebees. In valley, near Ansob Pass, Tajikistan. April 2014
Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK
Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2
Porcelain orchid (Chloraea magellanica) with osmophores on lip fluorescing in UV light. Chile. Controlled conditions, focus stacked. Sequence 2/2
Chinese meadow-rue (Thalictrum delavayi) flowers. Male phase below with stigmas hanging below immature stigmas, female phase above after stamens have fallen
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
Cup fungus (Cookeina sp) growing on decaying wood on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Snowy owl (Bubo scandiacus) Female taking off the winter tree, Quebec, Canada
Man using pole to propel bamboo raft on Lijang / Li River, Karst peaks beyond. At dawn, Lijiang River Scenic Zone, Yangshuo county, Guangxi, China
Golden-collared Tanager (Iridosornis jelskii), in the Peruvian cloud forest. Wayqecha Biological Research Station, Peru. April. Cropped
Lapwing (Vanellus vanellus) Titchwell RSPB Reserve, Norfolk, England, UK. November
Red squirrel (Sciurus vulgaris) foraging in forest, climbing down tree trunk, Cairngorms National Park, Scotland, October
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015