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Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
Arabian Oryx (Oryx leucoryx) at sunset. Previously extinct in the wild, their conservation status is now stable thanks to conservation efforts.This male is part of a reintroduction program in Dubai
Toco Toucan (Ramphastos toco) head and beak profile portrait, Brazil
RF - Coquerels sifaka (Propithecus coquereli) jumping, Anjajavy Private Reserve, north west Madagascar
Great hammerhead shark (Sphyrna mokarran) in shallow water. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August
Common frog (Rana temporaria) portrait, Broxwater, Cornwall, UK. June
Small tortoiseshell butterfly (Aglais urticae) resting on old painted door. Dorset, UK, August
Black-headed gull (Chroicocephalus ridibundus) alighting on water at dawn, Cheshire, UK, November. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Burrowing owl (Athene cunicularia) fledgling with head on side, Pantanal, Brazil. November
Luminescent jellyfish / Mauve stinger (Pelagia noctiluca) in open water, Gozo Island, Malta. Mediterranean Sea
Lone tree in a barley field. Picardy, France
Silver Birch (Betula verrucosa) reflected in Loch Beinn a Mheadhoin. Glen Affric National Nature Reserve, Scotland, December
Mute Swan (Cygnus olor) with its beak in water. Fife, Scotland, November
Polar bear (Ursus maritimus) female and her two cubs on large iceberg off Baffin Island. Nunavut, Canada, August 2010
Queen Marys Tree, a veteran Sweet chestnut (Castanea sativa) over 600 years old, Cumbernauld, Scotland, UK, July
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
House Mouse (Mus musculus) peering through hole in cheese. Captive
Arctic tern (Sterna paradisaea) in flight with fish in beak for feeding chick, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, July
Three Strawberry poison frogs (Oophaga pumilio) on rock, Sarapiqui, Heredia, Costa Rica
Sword billed hummingbird (Ensifera ensifera) in flight, Guango, Napo, Ecuador
Group of African elephants (Loxodonta africana) silhouetted at sunrise, Okavango Delta, Botswana
Two Atlantic spotted dolphins (Stenella frontalis) over a shallow sand bank. On the left is a juvenile which does not have the characteristic spots of the adults
Humpback whale (Megaptera novaeangliae), fluke of female with male calf playing below, Vava'u, Tonga, Pacific Ocean
Tufted duck (Aythya fuligula) on water on a misty morning, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. June
Female Cheetah (Acinonyx jubatus) standing in long grass on the savannah, Okavango Delta, Botswana, Africa
Blackbird (Turdus merula) and falling snow. Leicestershire, UK, February
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Caribbean flamingo (Phoenicopterus ruber) flying, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Little pied cormorant (Microcarbo melanoleucos) perched on branch protruding out of water. Eden NSW, Australia
RF - Reindeer (Rangifer tarandus) walking up slope in snow. Svalbard, Norway, April. (This image may be licensed either as rights managed or royalty free.)
RF - Arctic fox (Alopex lagopus) in winter pelage, camouflaged in snow. Svalbard, Norway, April (This image may be licensed either as rights managed or royalty free.)
Goldeneye (Bucephala clangula) male on loch on misty morning, Loch Vaa, Cairngorms National Park, Scotland, UK, April
Kingfisher (Alcedo atthis) perched on branch, Black Forest, Baden-Wurttemberg, Germany
Iberian lynx (lynx pardinus) in distance, silhouetted on horizon, Sierra de Andujar NP, Andalusia, Spain. December
African elephant (Loxodonta africana) herd with calf, silhouetted, Botswana
White throated kingfisher (Halcyon smyrnensis) perched on a reed, Sunderban tiger reserve, West Bengal, India
Alpine ibex (Capra ibex) male in deep snow on a ridge during heavy snowfall, Gran Paradiso National Park, the Alps, Italy
Elephant (Loxodonta africana) female and calf under tree in the rain, Masai-Mara game reserve, Kenya
RF - European goldfinch (Carduelis carduelis) perched in field of Sunflowers (Helianthus annuus), Cadiz, Andalusia, Spain. July. (This image may be licensed either as rights managed or royalty free.)
Yellow wagtail (Motacilla flava) perched among flowering Rapeseed (Brassica napus), Scotland, UK. May
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Stink bug (Graphosoma lineatum). The Netherlands. September
Scarlet Macaw (Ara macao) couple in flight and breaking to land Corcovado National Park, Osa peninsula, Costa Rica
Large whip goby (Bryaninops amplus) with a parasitic copepod (with two trailing egg ribbons) perched on Red whip coral. Whale Rock. Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia
Yellow mongoose (Cynictis penicillata), two pups nose to nose. Kalahari Gemsbok National Park, Kgalagadi Transfrontier National Park, Northern Cape, South Africa
RF - Chinstrap penguins (Pygoscelis antarctica) displaying, Half-Moon Island, South Shetland Islands, Antarctica (This image may be licensed either as rights managed or royalty free.)
Roe deer (Capreolus capreolus) on a misty morning at the edge of a pond. Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. December