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Mute swan (Cygnus olor) on river in evening light, Gelderse Poort, near Nijmegen, The Netherlands. June
Egyptian goose (Alopochen aegyptiaca) in flight, London, UK, February
Goldeneye (Bucephala clangula) male and two females on woodland loch at dawn on misty morning, Loch Vaa, Cairngorms National Park, Scotland, UK, April
RF- Whooper Swan (Cygnus cygnus) standing on frozen loch at dawn in winter. Loch Insh, Cairngorms National Park, Scotland, UK
Caribbean flamingo (Phoenicopterus ruber) landing, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Lemon-yellow tree frog (Hyla savignyi) peering out from behind plant. April
RF - Adelie penguins (Pygoscelis adeliae) on blue ice berg. Graham Passage, Antarctic Peninsula, Antarctica. (This image may be licensed either as rights managed or royalty free.)
Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
Grey wolf (Canis lupus) standing among trees in snow-covered forest, Finland. November
Caribbean flamingo (Phoenicopterus ruber) feeding in waves, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August. Bookplate
RF - Atlantic puffin (Fratercula arctica) among sea campion (Silene maritima), Skomer Island, Pembrokeshire, UK. May
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Caribbean flamingos (Phoenicopterus ruber) line of adults standing in water, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
Broadhaven Beach, Pembrokeshire Coast National Park, Pembrokeshire, Wales, March 2012
Caribbean flamingo (Phoenicopterus ruber) flock in distance at dusk, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, August
Caribbean flamingos (Phoenicopterus ruber) silhouetted at dawn, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August
Puffin (Fratercula arctica) through soft focus vegetation. Great Saltee, Saltee Islands, County Wexford, Ireland, June
Bee orchid (Ophrys apifera), Powerstock Common, Dorset Wildlife Trust, Dorset, England, UK. June
Red fox (Vulpes vulpes) rear view sitting on grass close to its den, Derbyshire, UK
Dusky shark (Carcharhinus obscurus) swimming through light rays during sardine run. Wild Coast, Eastern Cape, South Africa. June
Maned wolf (Chrysocyon brachyurus) setting out to hunt in grassland at dusk. Serra da Canastra National Park, Brazil
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Snakelocks anemone (Anemonia viridis) growing on a stalk of Eelgrass (Zostera marina) on the seabed, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Female scalefin anthias (Pseudanthias squamipinnis) swimming in front of a school of glassfish (Apogonidae), whose shiny bodies are rendered into out of focus bokeh patterns. Laamu Atoll, Maldives
Mute swan (Cygnus olor) swimming on sun-dappled water, Lower Silesia, Poland
Mandarin duck (Aix galericulata) drake standing on frozen pond during snowstorm. Richmond Park, London, UK. January
Male Black woodpecker (Dryocopus martius) in snow, showing red crown. Kuusamo Finland December. Overall winner of the Swiss Vogelwarte photo competition 2021
Pygmy leaf-folding frog (Afrixalus brachycnemis) sitting on a fern frond, Tanzania, Africa
Song thrush (Turdus philomelos) looking for worms in a garden, Broxwater, Cornwall, UK. April
Bearded vulture (Gypaetus barbatus) flying, Ordesa National Park, Pyrenees, Spain
Thorny devil stick insect (Eurycantha calcarata), Willaumez Peninsula, New Britain, Papua New Guinea, December
Grey triggerfish (Balistes capriscus) portrait, Tenerife, Canary Islands. August
Portrait of a rescue dog on white background, one ear perked up
Buzzard (Buteo buteo) landing on branch, winter, Lorraine, France, February
Rosemary beetle (Chrysolina americana) feeding on Lavender (Lavandula angustifolia) in garden Cheshire, UK, June
RF - Sailing boat in mist at sunrise, reflected in Wimbleball Lake. Exmoor National Park, Somerset, England, UK. September 2020. (This image may be licensed either as rights managed or royalty free.)
RF - Small tortoiseshell butterfly (Aglais urticae) nectaring on Verbena (Verbena sp). Cornwall, England, UK. September. (This image may be licensed either as rights managed or royalty free.)
Scoophead shark (Sphyrna media), released specimen. Isla Chepillo, Rio Bayano, Panama
Cuban dogfish (Squalus cubensis). Exuma Sound, Bahamas
Atlantic sixgill shark (Hexanchus vitulus) swimming with open mouth. Cape Eleuthera, Bahamas
Blotched blue-tongue skink (Tiliqua nigrolutea) female gaping with tongue out, in defense against perceived threat. Lake Tantangara, Snowy Mountains, New South Wales, Australia
Mute swan (Cygnus olor) stretching its wings after preening on a misty morning. Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Arctic fox (Vulpes lagopus), blue colour morph in winter coat walking through snow. Hornstrandir Nature Reserve, Iceland. February
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
RF - Red-and-green macaws (Ara chloropterus) two with one taking off, Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
