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Ptarmigan (Lagopus muta) male on snowy ridge on Cairngorm mountain, Cairngorms National Park, Scotland, UK, March
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Raven (Corvus corax) calling in the snow, Kemijarvi, Finland, February
Sword-billed hummingbird (Ensifera ensifera) hovering in flight, North-Ecuador
Mountain hare (Lepus timidus) adult resting in snow hole, Scotland, UK, February
Red deer (Cervus elaphus) male stag roaring portrait, rut season, Jura, Scotland, UK, September
Golden working cocker spaniel puppy amongst bluebells in beech woodland, Micheldever Woods, Hampshire, UK
Cow killer velvet ant (Dasymutilla occidentalis) on white background, Tuscaloosa County, Alabama, USA September
Baby rabbit with a Roborovski Hamster (Phodopus roborovskii) sitting on its head
Oak tree (Quercus robur) silhouetted against night sky with stars, Brecon Beacons National Park, Wales, UK. December
Cavalier King Charles Spaniel, bitch with black-and-tan coat
Female Black-footed cat (Felis nigripes) in the desert at night, Karoo, South Africa. Taken on location for the BBC series Big Cats
Great crested grebe (Podiceps cristatus) adult with young on the back in the first morning light Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. May
Wood pigeon (Columba palumbus) flying with nesting material in beak, Lorraine, France, April
Deepsea jellyfish (Periphylla periphylla) juvenile, , Trondheims fjord, Norway, Atlantic ocean
Rabbit (Oryctolagus cunniculus) juvenile grooming ears, Wiltshire, UK, May. Sequence 1 of 3
Pallass cat (Otocolobus manul) licking lips whilst running over ice. East Mongolia. February
Sparrowhawk (Accipiter nisus) adult female, Norway, October
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
Short-finned pilot whale (Globicephala macrorhynchus) calf with fetal lines characteristic of newborn specimens, with pod. Tenerife, Canary Islands
Arctic fox (Alopex lagopus) camouflaged in winter pelage. Svalbard, Norway, April
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Geraldton wax flower (Chamelaucium uncinatum), nectar fluorescing in UV light. Western Australia. Controlled conditions, focus stacked. Series 1/2
Flame lily (Gloriosa superba), timelapse sequence from opening bud to flowering, tepals becoming reflexed. Flower pollinated by butterflies. Controlled conditions
Hummingbird hawk-moth (Macroglossum stellatarum) nectaring on Loofah (Luffa sp) flower. Xinjiang, China
Chinese sturgeon (Acipenser sinensis), captive in aquarium. Adults are no longer able to swim upriver to spawn due to Gezhouba and Three Gorges dams
Bubble eye goldfish (Carassius auratus) with upward pointing eyes and two large fluid-filled sacs.. In captivity, Shanghai Zoo, Shanghai, China
Bruising webcap mushroom (Cortinarius purpurascens) Annagarriff Wood NNR, Peatlands Park, County Armagh, Northern Ireland. November
Rock Ptarmigan (Lagopus muta) portrait, Inari, Finland, September
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) in flight, about to land. Ecuador, May
Pelagic thresher shark (Alopias pelagicus) being cleaned by a bluestreak cleaner wrasse (Labroides dimidiatus) at Monad Shoal off Malapascua Island, Philippines
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August
Mountain hare (Lepus timidus) in summer pelage amongst heather, Cairngorms National Park, Scotland, UK, August
Sparkling Violet-ear (Colibri coruscans) hummingbird in flight, Mindo cloud forest ecosystem, Ecuador, July
Golden eagle (Aquila chrysaetos) in flight over snow. Kalvtrask, Vasterbotten, Lapland, Sweden. January
Black working cocker spaniel in bluebells in beech woodland, Micheldever Woods, Hampshire, UK
California quail (Callipepla californica), male emerging from behind snowbank in winter, Mono Lake Basin, California, USA
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
RF- North American river otter (Lutra canadensis) captive, occurs in North America
Rocks and coastal rainforest. Cocos Island National Park, Costa Rica. 2018
Silky shark (Carcharhinus falciformis). Cocos Island National Park, Costa Rica
Parrot snake (Leptophis ahaetulla) in aggressive pose with mouth open. Costa Rica
Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica
Violet sabrewing hummingbird (Campylopterus hemileucurus) nectaring on Orchid. Costa Rica
Glowworm beetle (Phengodes sp. ) male flying, Tuscaloosa County, Alabama, USA Controlled conditionsGlowworm beetle (Phengodes sp.) male flying, Tuscaloosa County, Alabama, USA Controlled conditions. July
