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Red deer (Cervus elaphus) stag portrait, bellowing with bracken in antlers, rutting season, Bushy Park, London, UK, October
Two Fallow deer (Dama dama) bucks during fight, rutting season, Richmond Park, London, UK, October
Pair of Common cuttlefish (Sepia officinalis), female in foreground and male behind, note circular mating scars on female caused by the males suckers, Babbacombe Bay, Torbay, Devon, UK, May
Common blue damselfly {Enallagma cyathigerum}, mating pair, Little Bradley Ponds, Devon, UK. July. 2020VISION Book Plate
Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male in flight at lek, Cairngorms NP, Grampian, Scotland, UK, April
Red deer (Cervus elaphus) stag bellowing, Bradgate Park, Leicestershire, England, UK, October
Black grouse (Tetrao tetrix) male displaying at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Brown hare (Lepus europaeus) two males chasing female in snow covered field during a blizzard. Derbyshire, England, UK, February. Winner of British Seasons category of BWPA competition 2014
Moorland Clouded Yellow butterfly (Colias palaeno) male and female, Joutsa (formerly Leivonmaki), Finland, July
Silver-studded blue butterfly (Plebejus argus) pair mating, resting on a Pyrimidal orchid (Anacamptis pyramidalis) flower, Gwithian Towans, Cornwall, UK, July
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Great crested grebe (Podiceps cristatus) pair performing courtship display at dawn, backlit with surrounding by mist, Cheshire, UK, March
RF- Black Grouse (Lyrurus tetrix) displaying. Finland. March
Common cranes (Grus grus) displaying, Hula Valley, Northern Israel, January
Silhouette of two male Black grouse (Tetrao tetrix) displaying at lek at dawn, Cairngorms NP, Grampian, Scotland, UK, April. 2020VISION Book Plate
Male Black grouse (Tetrao tetrix) displaying at lek at dawn, Cairngorms NP, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male at lek, Creag Megaidh NNR, Highlands, Scotland, UK, April
Capercaillie (Tetrao urogallus) male displaying in pine forest in snow, Cairngorms NP, Highlands, Scotland, UK, January. Photographer quote
Portrait of pair of Bohar snappers (Lutjanus bohar) in their spawning aggregation. Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Indigo hamlets (Hypoplectrus indigo) pair spawning at dusk, George Town, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. Shark City, Ulong, Rock Islands, Palau
Capercaillie (Tetrao urogallus) male, displaying in pine forest, Cairngorms NP, Scotland, UK, April
Andean cock of the rock (Rupicola peruviana) male in courtship ritual, Andean Mountains, Merida, Venezuela
Wolf Spiders (Pardosa sp. ), male RIGHT waving palps in courtship displayWolf Spiders (Pardosa sp.), male RIGHT waving palps in courtship display. Peak District National Park, Derbyhsire, UK. May
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
Pair of Common toads (Bufo bufo) in amplexus underwater, Belgium, March
Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June
Male Indian peafowl (Pavo cristatus) displaying, captive
Couchs Spadefoot (Scaphiopus couchii) adult at night calling with vocal sac inflated. Laredo, Webb County, South Texas, USA, April
Atlantic salmon (Salmo salar) moving upriver to spawn. Lligwy River nr Betws y Coed, Gwynedd, North Wales, UK, November
African lion (Panthero leo) male giving flehmen grimace, Tanzania, East Africa
European hare (Lepus europaeus) mating pair boxing in field, Slovakia
Pair of King Penguins (Aptenodytes patagonicus) on Gold Beach, South Georgia Island, Sub Antarctica
King penguin courtship (Aptenodytes patagonicus) Falkland Islands
Great crested grebes (Podiceps cristatus) performing their elaborate courtship ritual at dawn, Cheshire, UK November
