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Skeleton shrimp (Caprella septentrionalis) mating Novaya Zemlya, Russian Arctic, July
Western capercaillie (Tetrao urogallus) male calling at lek, with female, Tver, Russia. May
Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April
Gunnison sage-grouse (Centrocercus minimus) male displaying at a lek. Gunnison County, Colorado. April
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Great Egret (Casmerodias / Ardea alba) courtship display, backlit, Orlando, Florida, USA
European bee-eater (Merops apiaster) pair in courtship display, Bulgaria, May 2008
Three European Hare (Lepus europaeus) chasing, a courtship behaviour. Wales, UK, June
Eurasian dotterel (Charadrius morinellus) pair mating in breeding habitat on upland plateau of Grampian mountains, Cairngorms NP, Scotland, UK, May 2011
Great crested grebe (Podiceps cristatus) pair of adults during part of their elaborate courtship ritual during the very last few rays of evening sunlight, Derbyshire, UK, March
Black grouse (Tetrao tetrix) two males displaying and fighting at lek, Cairngorms NP, Grampian, Scotland, UK, April
Rear view of Black grouse (Tetrao tetrix) male displaying at lek, Cairngorms National Park, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male displaying at lek, Cairngorms National Park, Grampian, Scotland, UK, April
Red deer (Cervus elaphus) stag bellowing during autumn rut, Richmond Park, London, England, UK, October
Red deer (Cervus elaphus) young stag tasting the air (flehmen response) Bradgate Park, Leicestershire, England, UK, October
Whooper swan (Cygnus cygnus) pair on loch in heart shape, Cairngorms National Park, Scotland, UK, November
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
Common dolphins (Delphinus delphis) pair in mating ritual, Atlantic ocean, Portugal, July
Woodcock orchid (Ophrys cornuta / scolopax) with mating Black veined white butterfliesWoodcock orchid (Ophrys cornuta/scolopax) with mating Black veined white butterflies (Aporia crataegi), Bela Reka, Eastern Rhodope Mountains, Bulgaria, May 2013
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Five-spot burnet moths {Zygaena trifolii} mating on Pyramidal orchid {Anacamptis pyramidalis} Cornwall, UK
Satellite male Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May
Strawberry poison frog (Oophaga pumilio) pair mating, Sarapiqui, Heredia, Costa Rica
Lesser kestrel (Falco naumanni) male flying past during courtship with Mole cricket in its talons. Extremadura, Spain
Yellow-flecked glassfrogs (Sachatamia albomaculata) mating pair, Osa Peninsula, Costa Rica
Four-spot orb weaver spider (Araneus quadratus) male approaching larger female to mate. Peak District National Park, Derbyshire, UK. October
Emerald damselflies (Lestes sponsa) mating, with male above. Peak District National Park, Derbyshire, UK. September
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Red deer stag (Cervus elaphus) bellowing in rutting season in morning with steam rising from its body, New Forest, Hampshire, UK
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Two male Common cuttlefish (Sepia officinalis) compete for a female (on right) during courtship, Torbay, Devon, UK, May. 2020VISION Exhibition. 2020VISION Book Plate
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Common toads (Bufo bufo) in mating ball underwater, Belgium, March
Lemur leaf frog (Hylomantis lemur) mating pair, captive, Critically endangered, from Central America. Please credit Michael Kern / Zoo Atlanta
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Pipevine Swallowtail Butterfly (Battus philenor) pair mating. Sinton, Corpus Christi, Coastal Bend, Texas, USA, June
Male Andean Cock-of-the-Rock (Rupicola peruvianus) displaying. Cock-of-the-Rock Lodge, Peru, August
Goldies Bird of Paradise (Paradisaea decora) male displaying to female at lek, Fergusson Island, Milne Bay Province, Papua New Guinea. Near-threatened
Mating pair of King Penguins (Aptenodytes patagonicus) on Gold Beach, South Georgia Island, Sub Antarctica
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
