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White stork (Ciconia ciconia) pair displaying on their nest built on the chimneys of Knepp Castle, Sussex, UK, June
Northern gannets (Morus bassanus) pair displaying amongst campion, Bempton Cliffs, Yorkshire, UK, May
Pair of mating Big blue octopuses (Octopus cyanea) where the male, on the right, is fertilising the female using its fertilising tentacle. Indian ocean, Maldives
Gentoo penguins (Pygoscelis papua) pair, Sea Lion Island, Falkland Islands
Black-browed albatross (Thalassarche melanophris) pair in courtship. Falkland Islands
Waved albatross (Phoebastria irrorata), pair mutual preening in courtship, Espanola Island, Galapagos Islands
Kingfisher (Alcedo atthis) male passing fish to female, courtship behaviour, Lorraine, France, March
Pink salmon (Oncorhynchus gorbuscha) migrate up river, Vancouver Island, Canada
Bilingual frog (Crinia bilingua) adult male calling, Adelaide River, Northern Territory, Australia, January
Stag beetle (Lucanus cervus) males fighting at dusk in oak wood, the Netherlands, June
Two Black-browed albatross (Thalassarche melanophris) rub their bills together - part of a bonding ritual. West Point Island, The Falkland Islands
Aerial view of Lesser flamingos (Phoeniconaias minor), mating display Lake Magadi, Rift Valley, Kenya
Arctic tern (Sterna paradisaea) pair mating, on rocks on beach. Hailuoto, Finland. May
Red deer (Cervus elaphus) stags fighting during the rutting season, Bushy Park, London, UK October
White-lipped deer (Cervus albirostris) walking on the Tibetan plateau, Serxu, Garze Prefecture, Sichuan Province, China
Clearfin lionfish (Pterois radiata) mating. The female (left) has just released a raft of eggs, which is visible between her tail and the red face of the male
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
African grey hornbill (Lophoceros nasutus) calling in display, perched on tree snag. Janjanbureh, Gambia
Azure / common coenagrion damselflies (Coenagrion puella) pair mating, Meeth Quarry, Devon, UK. July
Sockeye salmon (Oncorhynchus nerka) migration, Adams River, British Columbia, Canada. October
Galapagos hawk (Buteo galapagoensis) landing on mating pair of Galapagos giant tortoise (Chelonoidis vandenburgh) Alcedo Volcano, Isabela Island, Galapagos
Waved albatrosses (Phoebastria irrorata) courting, Punta Suarez, Espanola Island, Galapagos
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Common Merganser (Mergus merganser) female with ducklings, in threat posture at the approach of a second female, Lansing, New York, USA
Leopard (Panthera pardus) pair mating behaviour, Okavango Delta, Botswana
Black Grouse (Lyrurus tetrix) fighting on icem Finland. March
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Eurasian rollers (Coracias garrulus) courting on branch, Hungary. June
Small emerald spreadwing (Lestes virens) pair mating. Klein Schietveld, Brasschaat, Belgium. August
Blue-tailed damselfly (Ischnura elegans) pair mating on stem. Brasschaat, Belgium. August
King cormorants (Phalacrocorax atriceps) in courtship behaviour at their nest site. Saunders Island, Falkland Islands
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Pair of Western clown anemonefish (Amphiprion ocellaris) spawning orange eggs on the rock beneath their Magnificent sea anemone (Heteractis magnifica) home on a coral reef
Fallow deer (Dama dama) herd at dawn, Holkham, Norfolk, England, UK, October
Red deer (Cervus elaphus) stag roaring during the rut, Leicestershire, UK, October. Non-ex
Red deer {Cervus elaphus} adult male in autumnal bracken during the October rut. Leicestershire, UK
Silhouette of Red Deer (Cervus elaphus) stag calling during rut, Dyrehaven, Denmark
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Jaguars (Panthera onca) courting pair on a sand bank. Cuiaba River, Northern Pantanal, Mato Grosso, Brazil
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Magellanic penguin (Spheniscus magellanicus) calling at the entrance of burrow, courtship display, Carcass Island, Falkland Islands, October
Clarion Lizard (Urosaurus clarionensis) displaying, IUCN Vulnerable, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Common toad (Bufo bufo) pair in amplexus, Deeside, Scotland April