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RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
Pair of mating soldier beetles (Rhagonycha fulva) on Common ragwort (Senecio jacobaea), Arne RSPB reserve, Dorset, England, UK, August
Red deer (Cervus elaphus) stags fighting during rut, Richmond Park, London, England, UK, October
Red deer (Cervus elaphus) stag chasing another during a fight, rutting season, Bushy Park, London, UK, October
Two Red deer (Cervus elaphus) stags fighting, rutting season, Bushy Park, London, UK, October
Capercailie (Tetrao urogallus) male displaying from branch of Scots pine tree. Cairngorms National Park, Highlands, Scotland, UK, March
Common goldeneye (Bucephala clangula) male performing its mating display. Southern Norway. March
Spotted owlet (Athene brama) pair on a branch, Bharatpur / Keoladeo Ghana National Park, India
Red deer (Cervus elaphus) stag bellowing during rut, at night with full moon, Derbyshire, UK
Red deer (Cervus elaphus) stag bellowing in mist at sunrise, rutting season, Bushy Park, London, UK, October
Red deer (Cervus elaphus) stag thrashing bracken, rutting season, Bushy Park, London, UK, October
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
Red deer (Cervus elaphus) stag roaring at dusk during rutting season, Cheshire, UK October
Guianan Cock-of-the-rock (Rupicola rupicola), during courtship display, Guianan Shield, Suriname
Argus pheasant / Great argus (Argusianus argus) male displaying, Danum Valley forest reserve, Sabah, Borneo, Malaysia
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Mandarinfish (Synchiropus splendidus) pair spawning. The larger male (behind) is lifting the female into the spawning rise on his pectoral fin, the smaller female (foreground) is releasing eggs
Male Carolas Parotia (Parotia carolae) bird of paradise displaying on his display court to five females perched above, Crater Mountain Wildlife Management Area, Eastern Highlands Province
Waved Albatross (Phoebastria irrorata) courtship, Punta Cevallos, Espaola Island, Galapagos Islands, Ecuador, South America. Endemic, Critically endangered
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Red deer (Cervus elaphus) stag thrashing bracken, rutting season, Bushy Park, London, UK, October. 2020VISION Book Plate
Humpback whale (Megaptera novaeangliae australis) courtship, with the female above showing her abdomen and the male below. Vava'u, Tonga. Pacific Ocean
Kori bustard (Ardeotis kori) male, performing mating display, silhouetted at dusk, El Karama Wilderness Conservancy, northern Kenya
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Male Common peafowl (Pavo cristatus) displaying feathers, Ranthambore NP, Rajasthan, India
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA. When displaying
Tench (Tinca tinca) during spawning period amongst European watermilfoil (Myriophyllum spicatum), Sion, Valais canton, southwestern Switzerland
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
Kottigehar dancing frog (Micrixalus kottigeharensis), male waving foot and calling, territorial behaviour. Agumbe, Western Ghats, India. Endemic
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October
Giant barrel sponge (Xestospongia testudinaria) releases a large cloud of gametes as it spawns on a coral reef, while a Threadfin hawkfish (Cirrhitichthys aprinus) looks
Sockeye salmon (Oncorhynchus nerka) males fighting in river Adams River, British Columbia, Canada. October
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
