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Indian peafowl (Pavo cristatus) peacock walking and displaying feathers, Yala National Park, Southern Province, Sri Lanka
RF - Arctic fox (Vulpes lagopus) male, blue colour morph. Hornstrandir Nature Reserve, Iceland. March
Wolf (Canis lupus) male on rocky snow covered slopes. Ulley Valley in the Himalayas, Ladakh, India
Red deer (Cervus elaphus) stag in autumn woodlands, Bradgate Park, Leicestershire, UK, October
RF - Red Deer stags (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
Great crested newt (Triturus cristatus) male in breeding condition, amongst Soft hornwort (Ceratophyllum submersum), Herefordshire, England, UK, May. Captive
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Red deer (Cervus elaphus) stag bellowing during rut, at night with full moon, Derbyshire, UK
European Adder (Vipera berus) portrait of a female and a male. Aberdeenshire, Scotland, May
Adult male Sparrowhawk (Accipiter nisus) with prey perched on a branch, Dumfries, Scotland, UK, November
Large male Napoleon wrasse (Cheilinus undulatus) yawning. Ras Mohammed, Sinai, Egypt. Red Sea
Male Yellow-headed jawfish (Opistognathus aurifrons) blows out the clutch of eggs he is incubating in his mouth to oxygenate them. East End, Grand Cayman, Cayman Islands, British West Indies
A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October
Male Great white shark (Carcharodon carcharias) with sunrays, Guadalupe Island, Mexico, Pacific Ocean
Orange Tip Butterfly (Anthocharis cardamines) male resting on Bluebell (Endymion / Hyacinthoides nonscriptus). Peak DIstrict National Park, Derbyshire, UK. April
Guianan Cock-of-the-rock (Rupicola rupicola), during courtship display, Guianan Shield, Suriname
American goldfinch (Carduelis tristis) male in breeding plumage, perched in Eastern redbud flowers in spring, New York, USA May
Magnificent bird of paradise (Diphyllodes magnificus) male, Lower montane highlands near Mount Hagen, Enga Province, Papua New Guinea. Endemic
Banded demoiselle (Calopteryx splendens) male close up detail of head and thorax, Hertfordshire, England, UK, June. Captive. Focus stacked
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Melanistic leopard / Black panther (Panthera pardus) on territorial patrol on track, Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Siberian musk deer (Moschus moschiferus) male, Irkutsk, Russia. March
Black woodpecker (Dryocopus martius) male on snowy tree trunk, Kuusamo, Finland, February
Pine grosbeak (Pinicola enucleator) male and female, Liminka, Finland, January
Feral Goat (Capra hircus) male, Inverness-shire, Scotland, July
Wild Boar (Sus scrofa) males fighting in snow. The Netherlands, January
Moose (Alces alces) head portrait of bull standing in forest clearing, Cap Breton Highlands National Park, Nova Scotia, Canada, September
Boston Terrier, head portrait of male
King Penguin (Aptenodytes patagonicus) adult surrounded by huddled chicks, riding out a snowstorm at Gold Harbor, South Georgia
Magyar Vizsla / Hungarian Pointer, head portrait of smooth coated, tan coloured male, sitting
Black howler monkeys (Alouatta caraya) male and two females calling from tree, captive, Apenheul Park, Netherlands
Close up face portrait of male Orang Utan (Pongo pygmaeus) Captive, Netherlands
Moose (Alces alces) bull portrait, Baxter State Park, Maine, USA
Grevy zebra (Equus grevyi) stallions fighting, Samburu National Reserve, Kenya
Lone male Bottlenose dolphin (Tursiops truncatus) in shallow water over seagrass. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Humpback whale (Megaptera novaeangliae), fluke of female with male calf playing below, Vava'u, Tonga, Pacific Ocean
Kori bustard (Ardeotis kori) male, performing mating display, silhouetted at dusk, El Karama Wilderness Conservancy, northern Kenya
Iberian ibex (Capra pyrenaica), adult male, walking along mountain ridge. Sierra Nevada National Park, Andalusia, Spain. June
Goldeneye (Bucephala clangula) male on loch on misty morning, Loch Vaa, Cairngorms National Park, Scotland, UK, April
Silhouette of Red deer (Cervus elaphus) stag standing under a tree, Bradgate Park, Leicestershire, UK. October
Banded demoiselle dragonfly (Calopteryx splendens) male and female silhouetted, resting on reeds. Lower Tamar Lakes, Cornwall, England, UK. June
Kudu (Tragelaphus strepsiceros) male silhouetted, Kruger National Park, South Africa. Digital silhouette
Alpine ibex (Capra ibex) male in deep snow on a ridge during heavy snowfall, Gran Paradiso National Park, the Alps, Italy
RF - Puffins (Fratercula arctica) pair, resting on cliffside among flowering Sea thrift (Armeria maritima), Great Saltee Island, County Wexford, Republic of Ireland. May
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Japanese dwarf flying squirrel (Pteromys volans orii) male sitting in tree, portrait. Hokkaido, Japan. March
Red deer (Cervus elaphus) stags running in herd across moorland, February
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
