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Sperm whales (Physeter macrocephalus) aerial shot showing much larger male on right, with female nursery group. Baja California, Mexico
Vermillion Waxcap (Hygrocybe miniata) Group of three toadstools, Buckinghamshire, England, UK, October - Focus Stacked
Himalayan ibex (Capra sibirica) during a snowfall, coming down the mountain at dusk to sleep at lower altitude, around Kibber village, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains
Shoal of Piraputanga, (Brycon hilarii), close to the surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia
Shrimpfish / Razorfish (Aeoliscus punctatus / strigatus) have adapted to a vertical posture and swim with their head downward. Lembeh, North Sulawesi, Indonesia
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Red-and-green macaws (Ara chloropterus) perched and in flight over Buraco das Araras, the Sinkhole of the Macaws, Jardim, Mato Grosso do Sul, Brazil. September
Aerial view of African elephants (Loxodonta africana) looking for the shade under trees on a very hot day, Northern Botswana
Plain / Burchells zebras in dust (Equus burchelli) Nxai Pan National Park, Botswana
Mexican free-tailed bats, (Tadarida brasiliensis), leaving Bracken Cave, Texas. Bracken Cave is the summertime home of over 15 milions bats
Brandts cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig
Great flamingo (Phoenicopterus roseus) feeding in sludge at bottom of river, Pont De Gau Park, Rhone River, France
Mountain hare (Lepus timidus) three adults on a snow slope high in the Cairngorm Mountains, Cairngorms National Park, Scotland, UK, February
White-throated Mountain Gem (Lampornis castaneoventris), Green-crowned Brilliant (Heliodoxa jacula) group of three with one in flight, Savegre, Costa Rica
Leaf cutter ants (Atta sp) carrying plant matter, Costa Rica
King penguin (Aptenodytes patagonicus) chick creche, Volunteer Point, East Falkland, Falkland Islands, October
Striated caracara (Phalcoboenus australis) group with one landing from above with only wings and tail visible. Carcass Island, Falkland Islands, October
King penguin (Aptenodytes patagonicus) colony, Volunteer Point, East Falkland, Falkland Islands, October
Zebra at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Blue morpho (Morpho helenor) butterflies feeding on mango fruit, butterfly observatory, La Paz Waterfall Gardens, Costa Rica
Whooper swans (Cygnus cygnus), flying at sunset, Caerlaverock Wildfowl & Wetland Trust, Dumfries & Galloway, Scotland, November 2016
Pronghorn antelope (Antilocapra americana) females, Saskatchewan, Canada
Shadowfin soldierfish (Myripristis adusta) schooling over coral reef. Dampier Strait, Raja Ampat, West Papua, Indonesia, March 2016
Colombian spider monkey (Ateles fusciceps rufiventris) group. Captive, Critically endangered species
South American sea lion (OtariaAflavescens) colony on beach, Punta Norte, Peninsula ValdesSouth American sea lion (Otariaflavescens) colony on beach, Punta Norte, Peninsula Valdes, Argentina
Swaynes hartebeest (Alcephalus busephalus swaynei), Senkelle Swaynes Hartebeest Sanctuary, Ethiopia
Emperor penguin (Aptenodytes forsteri), Chicks on ice, Snow Hill Island, Antarctic Peninsula
Elephant (Loxodonta africana) three calves walking down to Chobe River, Etosha National Park, Namibia, March
Northern gannets (Morus bassanus). Bass Rock, Scotland, UK. August
Mute swan (Cygnus olor) flock on snow, Hazerswoude, The Netherlands, February
Whooper swans (Cygnus cygnus) in misty hot springs, Lake Kussharo, Hokkaido, Japan, February
Atlantic Puffins (Fratercula arctica) in flight in mist, Skomer Island, Wales, UK, August
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Galapagos sea lions (Zalophus wollebaeki) swimming in the surf, Galapagos
Honey bees (Apis mellifera) forming living bridge, Kiel, Germany, June
Andean flamingos (Phoenicopterus andinus), Laguna Colorada, Bolivia. December 2016
Golden-crowned Sifakas or Tattersalls Sifakas (Propithecus tattersalli) feeding in forests near Andranotsimaty, Daraina, north east Madagascar. Critically Endangered
Red-crowned Cranes (Grus japonicus) flying at twilight, Hokkaido, Japan, February
Chamois (Rupicapra rupicapra) in deep snow trying to struggle their way out, Gran Paradiso National Park, Italy, November
Golden orb spiders (Nephila sp) on the shores of Lake Er Hai. Dali, Yunnan, China
Brown Hare (Lepus europaeus), Bayern, Germany. April
Yellowfronted canaries (Crithagra mozambica) at waterhole, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa. June
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
Dalmatian pelicans (Pelecanus crispus) squabbling over fish, Lake Kerkini, Greece, February
Japanese / Red-crowned crane (Grus japonicus) three coming into land, Hokkaido Japan February
Soft corals (Dendronephthya hemprichi) growing in very shallow water in the shade provided by a jetty, while a school of Silversides (Atherinomorus lacunosus) circle. Berenice Jetty, Aqaba, Jordan
