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Red mucronata (Balanophora harlandii), a parasite on woody roots with flowers resembling fungi. Wild Elephant Valley, Xishuangbanna, Yunnan, China
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Bar headed geese (Anser indicus), four in flight, one calling. Coming in to land at breeding site, Qinghai Lake, Qinghai Province, China
Painted lady butterflies (Vanessa cardui), three nectaring on potted Chrysanthemums (Chrysanthemum sp) in evening light. Urumqi Botanic Garden, Wushi, Xinjiang Uygur Autonomous Region, China
Ornamental koi carp (Cyprinus carpio) group with open mouths at lake surface. Xishuangbanna Tropical and Flowers Garden, Jinghong, Yunnan, China
Incarvillea (Incarvillea compacta) with raindrops on petals. Near Zhedou Shan Pass, Sichuan Province, China
Cremanthodium (Cremanthodium brunneopilosum). In wet meadow between Barkham and Ruo Er Gai, Sichuan Province, China
Mountain hare, (Lepus timidus), four animals on snowy hillside in winter, Scotland, UK.January
Red deer (Cervus elaphus) stags in snowy Pine forest. Scotland, UK. March
Dead leaf / Orange oak leaf (Kallima inachus) butterflies feeding on sap on tree trunk. Mimic leaves with wings closed. Wild Elephant Valley, Xishuangbanna, Yunnan Province, China
Crested ibis (Nipponia nippon) group roosting in tree at dusk. Captive bred. Yangxian, Shaanxi, China
African lion, (Panthera leo) confronted by a herd of African buffalo / Cape buffalo (Syncerus caffer), Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
RF - Pied kingfisher (Ceryle rudis) group of three perched on branch, squabbling, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Grayling (Thymallus thymallus) in the upper reaches of the Lena River. Baikalo-Lensky Reserve, Siberia, Russia. August 2018
Shoal of three Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka
Pod of Sperm whale socializing, (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean
Starlings (Sturnus vulgaris) resting on Blackpool Beach, UK. October
Emperor penguin (Aptenodytes forsteri) colony, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Brown noddies (Anous stolidus) perched on rock face, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador
Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April
Red deer (Cervus elaphus) herd of female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) stag bellowing, surrounded by hinds, Surrey, UK, September
Flock of Mute swans (Cygnus olor), Abbotsbury Swannery, Dorset, England, UK, September
Greylag goose pair (Anser anser) in flight over flooded pastureland with many grazing wildfowl, Gloucestershire, UK, September
Mountain hare, (Lepus timidus), four animals on snowy hillside in winter, Scotland, UK, February
Red squirrel, (Sciurus vulgaris), three animals backlit on pine branch, Cairngorms National Park, Scotland, UK.May
Grey seal (Halichoerus grypus) pup on a breeding beach, Orkney, Scotland, UK, April
Grey seal (Halichoerus grypus) huge group hauled out on a beach, Island of Mingulay, west coast of Scotland. May
Grey seal (Halichoerus grypus) using freshwater pools along the shore line and in the adjacent fields as place to learn to swim, Orkney, Scotland, UK, April
Close up of a colony of Jewel anemones (Corynactis viridis) Scotland, UK, October
White tent making bat (Ectophylla alba) roosting in Heleconia leaf, La Selva Field Station, Costa Rica
Family of six Mute swans (Cygnus olor) at sunrise, silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
Elegant anemone (Sagartia elegans) and Plumose anemone (Metridium senile) with fish swimming in plankton bloom in background, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
The Bass Rock home to large Northern gannet (Morus bassanus) Firth of Forth, Scotland, UK. August
Gannet (Morus bassanus) chick in a line of adults. Shetland Islands, Scotland, UK, August
RF - Mute swans (Cygnus olor) flock of non breeding adults, Fleet lagoon Dorset, England, UK, June
RF - Herring gull (Larus argentatus) with blue sky, England, UK, Digital composite
Sea otters (Enhydra lutris) resting on ice, Alaska, USA, June
Sperm whales (Physeter macrocephalus) aerial shot showing much larger male on right, with female nursery group. Baja California, Mexico
Vermillion Waxcap (Hygrocybe miniata) Group of three toadstools, Buckinghamshire, England, UK, October - Focus Stacked
Himalayan ibex (Capra sibirica) during a snowfall, coming down the mountain at dusk to sleep at lower altitude, around Kibber village, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains
Shoal of Piraputanga, (Brycon hilarii), close to the surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia
Shrimpfish / Razorfish (Aeoliscus punctatus / strigatus) have adapted to a vertical posture and swim with their head downward. Lembeh, North Sulawesi, Indonesia
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December