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Caribbean flamingo (Phoenicopterus ruber), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd at sunrise, Masai-Mara Game Reserve, Kenya. November
Group pf Puffins (Fratercula arctica) resting on cliffs of Heimaey Island, Westman Islands, Iceland. July
Three female Putorana snow sheep (Ovis nivicola borealis) climbing up rocky mountainside with lamb following, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Three Exmoor ponies (Equus ferus caballus), semi-feral native breed, in Exmoor National Park, Somerset / Devon, England. November
A bloom of Sea goosberries (Pleurobrachia pileus), Birsay, Shetland, Scotland, North Atlantic Ocean, UK
Juvenile Southern elephant seals (Mirounga leonina) practice fighting near a King penguin (Aptenodytes patagonicus) colony, Gold Harbour, South Georgia Island
A school of Shoulderbar soldierfish (Myripristis kuntee) swimming close to the reef for prtection during the day, Hawaii, Pacific Ocean
Hundreds of Periodical cicada (Magicicada sp. ) nymphs ascending a tree trunk to metmorphose, Princeton, New JerseyHundreds of Periodical cicada (Magicicada sp.) nymphs ascending a tree trunk to metmorphose, Princeton, New Jersey, USA. June
Drone aerial view of group of African forest elephants (Loxodonta cyclotis) searching for mineral salts in Dzanga-Sangha National Park, Central African Republic
Alpha male Grey wolf (Canis lupus) biting angry young wolf to assert dominance, Finland. July
Four male Silver-washed fritillary butterflly (Argynnis paphia) aggregating on wildflower, Finland. July
Flock of Sanderlings (Calidris Alba) in flight with wind turbines in background, Atlantic Coast, Vendee, France. December
Blackbar soldierfish (Myripristis jacobus) sheltering inside small cave, emerging from it at night to forage for food, Scotts Head, Dominica; Eastern Caribbean
A huge school of Pacific cownose / Golden cownose rays (Rhinoptera steindachneri), Baja California, Sea of Cortez, Mexico
Caribbean flamingo (Phoenicopterus ruber) landing, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January
Caribbean flamingo (Phoenicopterus ruber) flock, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January
Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp)Swarm of Asian weaver ants (Oecophylla smaragdina) attacking a Finger-print ant (Diacamma sp.), Kubah National Park, Sarawak, Borneo
Wolf (Canis lupus) pack huddling together in snow-covered forest, Sumava National Park, Bohemian Forest, Czech Republic. Captive
Small flock of migrating Black-tailed godwits (Limosa limosa) in flight against a bright blue sky, River Coquet Estuary, Northumberland, UK. September
RF - Eschmeyers scorpionfish (Rhinopias eschmeyeri) composite image showing different colour variations on black background. Lembeh Strait, North Sulawesi, Indonesia
Red knot (Calidris canutus) flock at high tide roost. Snettisham, Norfolk, UK. October
Three Lined seahorses (Hippocampus erectus), two male and one female, feeding on plankton in an alkaline saltwater pond, The Bahamas, Caribbean
Emperor penguin (Aptenodytes forsteri) group on ice floe, Ross Sea, Antarctica. January
Blue pincushion (Brunonia australis). Tasmania, Australia. December
Dominant male Chimpanzee (Pan troglodytes troglodytes) walking in the mangrove, with female carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Juvenile Mute swan (Cygnus olor) in early morning light, Valkenhorst Nature Reserve, The Netherlands, Europe. August
Three Grey partridge (Perdix perdix) walking in snow. Near Nijmegen, the Netherlands. February
Group of Grey partridge (Perdix perdix) huddled for warmth in snowy field. Near Nijmegen, the Netherlands. February
Common starling (Sturnus vulgaris) murmuration in between high voltage powerlines and electricity pylons, The Netherlands, Europe. February
School of Yellow-ribbon sweetlips (Plectorhinchus polytaenia) swimming with the current, Raja Ampat, Indonesia, Pacific Ocean
Spinifex pigeon (Geophaps plumifera) flock on stony desert floor, Diamantina River, Queensland, Australia
Galah (Eolophus roseicapillus) flock of birds flying over grassland, Diamantina River, Queensland, Australia
Group of Spectacled flying fox (Pteropus conspicillatus) roosting in daytime camp in rainforest, Atherton Tablelands, Queensland, Australia
Group of Grey-headed flying fox (Pteropus poliocephalus) roosting, Toowoomba, Queensland, Australia
Dominant male Chimpanzee (Pan troglodytes troglodytes) in the mangrove, with female and infant vocalising, Conkouati-Douli National Park, Republic of Congo, Africa
Green turtle (Chelonia mydas) visiting a cleaning station, Galapagos, South America, Pacific Ocean
Female Galapagos sea lion (Zalophus wollebaeki), sleeping together, likely to be related, Santa Fe Island, Galapagos, South America
Galapagos penguin (Spheniscus mendiculus) roosting on rocks in late afternoon, Isabela Island, Galapagos, South America
Eider ducks (Somateria mollissima) floating on waves, Iceland. March
Snow bunting (Plectrophenax nivalis) flock resting on a roof, Iceland. March
Iceland horses in winter, western Iceland. March
Aerial view of Caribbean flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
Caribbean flamingo (Phoenicopterus ruber) flock moving to other parts of the lagoon in search of food, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Caribbean flamingo (Phoenicopterus ruber), group, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Caribbean flamingo (Phoenicopterus ruber) flock, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Bookplate
Crescent-tail bigeye (Priacanthus hamrur) school on a coral reef, shot with a long exposure. North Male Atoll, Maldives. Indian Ocean
Adult Shags (Phalacrocorax aristotelis) with young, gathered on rocks, Treshnish Isles, Southern Hebrides, Scotland, UK, Atlantic Ocean. August
