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RF - Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Serengeti National Park, Tanzania. September
Nile crocodile (Crocodylus niloticus) waiting to attack White-bearded wildebeest (Connochaetes taurinus albojubatus) as they cross the Mara River
Coyote (Canis latrans) foraging in deep winter snow disturbed by grazing American bison (Bison bison) Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Smallmouth grunt (Haemulon chrysargyreum) fish swim over a coral reef, Cancun, Mexico. Slow shutter speed image
Laughing Gull (Larus atricilla) flock in flight, Sisal, Yucatan Peninsula, Mexico, January
Whooper Swans (Cygnus cygnus) in snow. Japan, February
Leucistic white lions (Panthera leo krugeri) male and juveniles, Captive
Three Marbled white butterflies (Melanargia galathea) nectaring on Greater knapweed flowers (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June
A school of filefifsh take shelter and feed on invertebrates (bottom of frame) found on sargassum in the Sargasso Sea
Blue tit (Cyanistes caeruleus) Sierra de Grazalema Natural Park. Southern Spain. July
Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site
Four Pygmy seahorses (Hippocampus bargibanti) on a Seafan / Gorgonian (Muricella sp. ) including one or several pregnant males. New Caledonia, Pacific Ocean
Crab-plover (Dromas ardeola) three flying, two with crabs in beaks. Oman, June
RF - Impala (Aepyceros melampus) at water with reflections, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
RF - Cape buffalo (Syncerus caffer) drinking at night, Zimanga private game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
Rocky mountain bighorn sheep (Ovis canadensis canadensis) grazing. Lamar Valley, Yellowstone National Park, Wyoming, USA. January
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Group of Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at dawn. Kasanka National Park, Zambia
RF- Young Ring-tailed lemurs (Lemur catta) carried on mother's back, Madagascar
African jackass penguins (Spheniscus demersus) three heading to sea in the early morning to forage for food, Cape Town, South Africa, Endangered IUCN
Atlantic walruses (Odobenus rosmarus) resting on ice, with two large individuals facing off just before the one on the right stabbed the other with his tusks, Svalbard, Norway, June
Southern right whales (Eubalaena australis) aerial view of three adullts engaged in social activity, photographed with the permission of the Department of Environmental Affairs, South Africa
Group of Sperm whales (Physter macrocephalus) family group swimming together, Indian Ocean, March
Aggregation of Sperm whales (Physeter macrocephalus) engaged in social activity. Indian Ocean, March
Large aggregation of Sperm whales (Physeter macrocephalus) engaged in social activity. Note the abundance of sloughed off skin in the water, resulting from tactile contact among the whales
Greater flamingo (Phoenicopterus roseus) flock, Pont Du Gau Park, Camargue, France
Springbok (Antidorcas marsupialis) herd on the move, Kgalagadi transfrontier park, South Africa
Female cheetah (Acinonyx jubatus) named Silgi (means bright future in Swahili) walking with 7 cubs. Masai Mara National Reserve, Kenya
Alcedo giant tortoise (Chelonoidis vandenburghi), group bathing. Alcedo Volcano, Isabela Island, Galapagos. 2017
RF - King penguin (Aptenodytes patagonicus), four returning to sea. St Andrews Bay, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
Guanacos (Lama guanicoe), five silhouetted on slope in evening. Torres del Paine National Park, Patagonia, Chile. December
Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt
Southern royal albatross (Diomedea epomophora) sub-adults, group displaying in tussock grassland. Campbell Island, New Zealand. February
Snow partridge (Lerwa lerwa) flock flying past snowy rockface, in evening light. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana
Rainbow lorikeet (Trichoglossus moluccanus) feeding chicks in nest hole. Lane Cove National Park, Sydney, New South Wales, Australia
South American Sealion, (Otaria byronia) breeding colony. Punta San Juan Reserve, (Reserva Nacional de Islas, Islotes y Puntas Guaneras) Peru
Bengal tiger (Panthera tigris) female and cubs swimming in pond. Ranthambore National Park, India
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Silky shark (Carcharhinus falciformis), three in Caribbean Sea off Gardens of the Queen National Park, Cuba
Water tulip (Cypella aquatilis) from above, in garden pond. Native to Brazil
Flowering rush (Butomus umbellatus) with stamens and stigma, nectar globules at base of carpels. Surrey, England, UK. Focus stacked
Afghanistan Juno iris (Iris cycloglossa) with yellow nectar guides. In cultivation, Kew Gardens, London, England, UK. Native to Afghanistan
Monarch butterflies (Danaus plexippus), three nectaring on Poinsettia (Euphorbia pulcherrima), resting on red bracts. In butterfly house, Xishuangbanna, Yunnan Province, China
Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California
Scarab beetles sleeping overnight in Persian marsh orchid (Dactylorhiza umbrosa). Turkey. June
Red mucronata (Balanophora harlandii), a parasite on woody roots with flowers resembling fungi. Wild Elephant Valley, Xishuangbanna, Yunnan, China
