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A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Panamic Cushion Sea Star (Pentaceraster cumingi) and a school of Yellow-tailed Grunt (Anisotremus interruptus), Galapagos Island, Ecuador. December
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Shoulderbar soldierfish (Myripristis kuntee) shoal. Molokini, Hawaii
Small sardine shoal inside the Gruta Azul dive site, eastern coast of Santa Maria Island, Azores, Portugal, Atlantic Ocean
Atlantic spotted dolphins, (Stenella frontalis), Formigas Islet dive site, Santa Maria Island, Azores, Portugal, Atlantic Ocean
Yellowfin goatfish (Mulloidichthys vanicolensis) shoal hovering over the reef, Hawaii
Coral reef scene, with Philippines chromis (Chromis scotochiloptera), hard coral (Acropora sp.) and purple elephant ear sponges (Ianthella basta)
RF - Maldive anemonefish (Amphiprion nigripes) swimming above their host Magnificent sea anemone (Heteractis magnifica) on coral reef
Giant coreopsis (coreopsis gigantea) flower, Zapato Islet, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, April
Pink footed geese (Anser brachyrhynchus) flock taking flight, near Udale Bay, Black Isle, Scotland, UK, October
European Bison (Bison bonasus) in winter, Białowieza National Park, Poland. January
Hermit crab (Dardanus arrosor), with Shoal of Ornate wrasse (Thalassoma pavo) Santa Maria Island, Azores, Portugal, Atlantic Ocean
Slime mould (Trichia botrytis) sporangia starting to split to release spores, on underside of rotting Oak (Quercus sp) log. Hertfordshire, England, UK. December. Focus stacked image
Bohemian waxwing (Bombycilla garrulus), flock at dusk with moon, Finland, February
Konik horses (Equus ferus caballus) interacting at sunset, floodplain of the river Rijn, Meinerswijk near Arnhem, the Netherlands. May
Vast swarm of Mayfly (Ephorom virgo) long exposure, emerged at the end of the summer Tudela, Navarra, Spain, September
White-lipped deer (Cervus albirostris) walking on the Tibetan plateau, Serxu, Garze Prefecture, Sichuan Province, China
Canada goose (Branta canadensis) flock taking off from a lake at sunrise. London, UK. November
Red stingray (Hemitrygon akajei) with Banded houndshark (Triakis scyllium) group and smaller fish in background. Chiba Prefecture, Honshu, Japan. September
Red stingray (Hemitrygon akajei), four in a row, viewed from below. Chiba Prefecture, Honshu, Japan. October
Leopard houndshark (Triakis semifasciata) group swimming amongst rays of light. San Diego, California, USA
Bronze whaler shark (Carcharhinus brachyurus) group feeding alongside Cape gannet (Morus capensis) during sardine run. Wild Coast, Eastern Cape, South Africa. June
Banded wobbegong shark (Orectolobus halei) resting on reef, Bullseye fish (Pempherididae) shoal in background. South West Rocks, New South Wales, Australia
Banded houndshark (Triakis scyllium) group above sea floor. Chiba Prefecture, Honshu, Japan. October
Atlantic mobula ray (Mobula hypostoma) group swimming in Caribbean Sea. Isla Mujeres, Quintana Roo, Mexico
King penguin (Aptenodytes patagonicus) breeding colony with chicks in creche, snow covered mountain in background. Andrews Bay, South Georgia. October 2017
Himalayan griffon vulture (Gyps himalayensis) Angsai Nature Reserve (Valley of the Cats), Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China
Blue sheep (Ovis nahoor) Angsai Nature Reserve (Valley of the Cats), Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China
Kiang or Tibetan Wild ass (Equus kiang) Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Dalmatian pelican (Pelicanus crispus) group squabbling over fish, Lake Kerkini, Greece
Green turtle (Chelonia mydas) resting on sea floor and two juvenile Golden trevally (Gnathanodon speciosus). Marsa El Shouna, Marsa Alam, Egypt
Green surf anemones (Anthopleura xanthogrammica) grow in shallow water below the temperate rain forest. Browning Pass, Port Hardy, Vancouver Island, British Columbia, Canada. North East Pacific Ocean
Common crane (Grus grus) flock feeding in field in snow, Champagne, France, February
Spotted eagle rays (Aetobatus narinari), in mangrove inlet, Elizabeth Bay, Isabela Island, Galapagos
Black-tailed godwit (Limosa limosa) flock in flight past Willow trees fringing a shallow lake, Gloucestershire, UK, February
A herd of wild konik horses running, with refection in water, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Highland cows, three silhouetted at dusk. Glengorm Estate, Isle of Mull, Scotland, UK. June
Yak (Bos grunniens) herd on plain beside river, mountains in background. Near Litang, Kham, Garze Tibetan Autonomous Prefecture, Sichuan, China. October 2016
Mealy Parrots (Amazona farinosa) feeding at the wall of a clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
King Penguins (Aptenodytes patagonicus) approached by an Antarctic Fur Seal (Arctocephalus gazella). A few pecks from the penguins caused the seal to retreat
Moths (Lepidoptera) swarming with flight trails, attracted by light during insect trapping and identification session. Long-term monitoring in De Kaaistoep Nature Reserve, Tilburg, The Netherlands
Moths (Lepidoptera) with flight trails, attracted to light during insect trapping and identification session. Long-term monitoring in De Kaaistoep Nature Reserve, Tilburg, The Netherlands. June
Dalmatian pelicans (Pelecanus crispus) squabbling over fish, Lake Kerkini, Greece. March
RF - Dalmatian pelicans (Pelecanus crispus) Lake Kerkini, Greece, March. This image may be licensed either as rights managed or royalty free.)
RF - North American river otters (Lontra canadensis) (probably female with two juveniles) on the frozen river edge.Yellowstone National Park, USA. January. Digitally stitched image
Monarch butterflies (Danaus plexippus) flying during a warm morning. When the overwintering period draws to an end large numbers become active. Michoacan, Mexico, February
European bison (Bison bonasus) herd in grassland. Eriksberg Wildlife and Nature Park, Blekinge, Sweden. June. Captive
