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Fallow deer heard (Dama dama) in heavy snow. Chatsworth Park, Derbyshire, Peak District National Park. January
Bewicks swans (Cygnus columbianus) small flock in flight, Gloucestershire, UK February
Longfin batfish (Platax teira) with Fusiliers (Caesio spp.) in the current in front of Baby Rock pinnacle. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea
Vibrant Red Sea reef scene, with orange female Scalefin anthias (Pseudanthias squamipinnis) swarming in front of Fire coral (Millepora dichotoma) feeding on plankton brought to the reef by currents
Green jacks (Caranx caballus) long exposure, La Paz, Mexico. Gulf of California, Sea of Cortez, East Pacific Ocean
Caribbean reef sharks (Carcharhinus perezi) swimming through the middle of a school of yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean
Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp)Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp.) and Scalefin anthias (Pseudanthias squamipinnis) and Magenta slender anthias (Luzonichthys waitei)
Long exposure of a juvenile Harlequin sweetlips (Plectorhinchus chaetodonoides), which dance as a defence from predators. The dance may mimic the movements of poisonous flatworms or may simply
Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. Shark City, Ulong, Rock Islands, Palau
California sea lion (Zalophus californianus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath Eureka oil rig, Los Angeles, California, USA, Pacific Ocean
School of Bohar snapper (Lutjanus bohar) swimming in formation along a coral reef. These fish are usually solitary and aggregate each summer in the Red Sea to spawn
Thomsons gazelle (Eudorcas thomsonii) herd with Wildebeest herd visible beyond, Masai-Mara game reserve, Kenya
Lyretail anthias / Goldies (Pseudanthias squamipinnis) over coral reef with soft corals (Dendronephthya sp) Egypt, Red Sea
Lesser flamingo (Phoeniconaias minor) flock, aerial view, Magadi lake, Kenya, September
Whale shark (Rhincodon typus) Galapagos Islands. Darwin Island and Arch, Galapagos
Blotcheye / Bigscale Soldierfish (Myripristis berndti), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
RF - Seafans (Melithaea sp. ) growing on dropoff, beneath schooling fish. Boo WindowsRF - Seafans (Melithaea sp.) growing on dropoff, beneath schooling fish. Boo Windows, Boo Islands, Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Three hairy sea spiders (Tanystylum grossifemorum) feeding on red soft coral (Gersemia rubiformis). Browning Pass, Port Hardy, Vancouver Island. British Columbia. Canada. North East Pacific Ocean
Hawksbill turtle (Eretmochelys imbricata) Port El Ghalib, Egypt, Red Sea
Herd of Wildebeest (Connochaetes taurinus) silhouetted at dusk, Serengeti NP, Tanzania, East Africa, January
Group of Manta rays (Manta birostris) feeding together on plankton in a shallow lagoon. Hanifaru Lagoon, Baa Atoll, Maldives. Indian Ocean
Caribbean reef shark (Carcharhinus perezi) swims over a coral reef. Grand Bahama Island, Bahamas. Tropical West Atlantic Ocean
A pod of Atlantic spotted dolphins (Stenella frontalis) riding on the bow wave of a boat. Sandy Ridge, Little Bahama Bank. Bahamas
Common / Eurasian cranes (Grus grus) small flock of juveniles taking flight over the frozen, snow covered pastureland on a foggy winter morning
Golden damselfish (Amblyglyphidodon aureus) swimming past coral reef wall with colourful soft corals. Bali, Indonesia
Flock of Cattle Egrets (Bubulcus ibis) in roosting trees at dusk, Imba Ndutu, Serengeti NP, Tanzania, East Africa
Large group of Californian sealions (Zalophus californianus) swimming underwater, Anacapa Island, California, Pacific Ocean
Bigscale soldierfish (Myripristis berndti), Sweetlips, Philippine chromis with Blueline snappers in background. Komodo National Park, Indonesia
Great dusky swift (Cypseloides senex) flock in front of Iguazu falls, Brazil / ArgentinaGreat dusky swift (Cypseloides senex) flock in front of Iguazu falls, Brazil/Argentina, from the Brazilian side. September
Sperm whale (Physeter macrocephalus) pod, Dominica, Caribbean Sea, Atlantic Ocean, January, Vulnerable species
School of Longfin spadefish (Platax teira), Kimbe Bay, West New Britain, Papua New Guinea
Mustang herd standing in snow. Black Hills Wild Horse Sanctuary, South Dakota, USA. January 2017
Bignose Unicornfish (Naso vlamingii), French Polynesia, Pacific Ocean
Orca whales (Orcinus orca), group of resident killer whales travelling together at dusk. British Columbia, Canada, Pacific Ocean
Dromedary camel (Camelus dromedarius) herd in a gorge with water on the Ennedi plateau. Ennedi Natural and Cultural Reserve, UNESCO World Heritage Site, Chad. September 2019
African elephant (Loxodonta africana) herd drinking at a waterhole, Mashatu Game Reserve, Botswana. June