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Northern gannets (Morus bassanus) breeding colony nesting on sea cliffs, Hermaness, Shetland, Scotland, UK. July
Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp. ) hunting a large school of Sardines (Sardina sp)Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex sp.) hunting a large school of Sardines (Sardina sp.), Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Group of California sea lions (Zalophus californianus) playing near the sea surface, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp)Brandt's cormorant (Phalacrocorax penicillatus) feeding on a bait ball of Sardines (Sardina sp.) being followed by a California sea lion (Zalophus californianus), Espiritu Santo Island
California sea lion (Zalophus californianus) close up with group playing in shallow water behind, Espiritu Santo Island, Baja California, Mexico, Sea of Cortez
Aerial view of massive arribada of Olive ridley turtle (Lepidochelys olivacea), with over 300, 000 females coming ashore to nest on 3 km of 15 km beach over period of three days and nights
Aerial view of thousands of female Olive ridley turtles (Lepidochelys olivacea) approaching beach during massive arribada, with over 300
Female Olive ridley turtles (Lepidochelys olivacea) coming ashore to nest while others leaves, at sunrise, during massive arribada. Playa Escobilla Sanctuary, Oaxaca, Mexico. Pacific Ocean
Lion-tailed macaque (Macaca silenus) infant, sitting among the troop with its finger in its mouth, Valparai, Tamil Nadu, India
Heermann's gull (Larus heermanni) breeding colony in flight and restless due to presence of pair of Peregrine falcon (Falco peregrinus)
Heermann's gulls (Larus heermanni) choosing and protecting nesting site. Rasa Island Special Biosphere Reserve, Sea of Cortez, Mexico. April
Springbok (Antidorcas marsupialis) herd in grassland after rain, Palmwag Conservancy, Damaraland, Namibia
Gemsbok (Oryx gazella) herd in desert after the wet season with Naukluft Mountains in the background, Namib-Naukluft National Park, Namibia
Herd of wild Onaqui horses (Equus ferus) trotting in dust. Onaqui Mountain Herd Management Area, Utah, USA. September
Two wild Onaqui stallions (Equus ferus) fighting for dominance in between their respective herd. Onaqui Mountain Herd Management Area, Utah, USA. September
Sika deer (Cervus nippon) herd feeding beside pool in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) herd walking past fallen tree in forest as one looks around, Land of the Leopard National Park, Russian Far East. Taken with remote camera. June
Shoal of Lowfin chub (Kyphosus vaigiensis) evading a Coral grouper (Cephalopholis miniata), Tubuai, French Polynesia, Pacific Ocean
Sulawesi black macaque / Celebes crested macaque (Macaca nigra) troop in forest, female with infant, Tangkoko National Park, northern Sulawesi, Indonesia. Critically endangered
Guanaco (Lama guanicoe) herd with the Towers granite rock formation and low cloud in background, Torres del Paine National Park, Patagonia, Chile
Puma (Puma concolor) female, stalking Guanaco (Lama guanicoe) herd in snow, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Bigeye trevallies (Caranx sexfasciatus) shoal. Darwin Island, Galapagos National Park, Pacific Ocean
Group of Spotted eagle rays (Aetobatus narinari) swimming in shallow water, Santa Cruz Island, Galapagos National Park, Pacific Ocean. Small repro only
Three Sri Lankan elephants (Elephas maximus maximus) with two calves walking along forest road, Yala National Park, Southern and Uva Provinces, Sri Lanka
Sally Lightfoot crabs (Grapsus grapsus) on rocks caught in the swell, Espanola Island, Galapagos National Park, Galapagos Islands
Herd of African elephants (Loxodonta africana) coming to drink at waterhole. Tsavo East National Park, Kenya
Pacific giant octopus (Enteroctopus dofleini) larvae close up, British Columbia, Canada, Pacific Ocean
Healthy coral reef landscape, Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
Common starlings (Sturnus vulgaris) large murmuration gathering before landing at winter roost, The Netherlands. Vogelwarte Photo Competition 2022 - Emotion category - Finalist
Aerial shot of a group of Mute swans (Cygnus olor) and gulls wintering together, Zurich, Switzerland. Vogelwarte Photo Competition 2022 - General Finalist
Gaggle of Canada goose (Branta canadensis) goslings on a lake, Burnhayes, Devon, UK, May
School of Scalefin anthias (Pseudanthias squamipinnis) females, swimming around Fire coral (Millepora dichotoma) feeding on plankton brought to the reef by currents, Ras Mohammed National Park
School of Burrito grunts (Anisotremus interruptus) huddling together in tight ball, Revillagigedo Islands, Mexico, Pacific Ocean
Shoal of large reef fish including Onespot snapper (Lutjanus monostigma), Oriental sweetlips (Plectorhinchus vittatus) and Sabre squirrelfish (Sargocentron spiniferum)
Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp)Large school of Glassfish (Apogonidae) swimming over colourful corals (Dendronephthya sp.) on a reef, Similan Islands, Thailand, Andaman Sea
Brent geese (Branta bernicla) small flock in flight over coastal feeding grounds, Lindisfarne National Nature Reserve, Northumberland, UK. February
Spotted deer (Axis axis), small herd leaping through grass, Bardia National Park, Terai, Nepal
Grey langurs (Semnopithecus entellus) family group, sitting and climbing in tree, Bardia National Park, Terai, Nepal
Spotted deer (Axis axis) herd in the forest, with just their antlers visible. Bardia National Park, Terai, Nepal
Leopard grouper (Mycteroperca rosacea) and Dentex (Dentex dentex) hunting a huge school of Sardines (Sardina pilchardus) in shallow water, Espiritu Santo archipelago, Sea of Cortez, Mexico
Horse-eye jack fish (Caranx latus) schooling around Caribbean reef sharks (Carcharhinus perezi) near Eleuthera Island, Bahamas, North Atlantic Ocean
Common dolphin pod (Delphinus delphis) jporposing, Horta island, Azores, Atlantic Ocean
Green sea turtle (Chelonia mydas) with shoals of damselfish (Pomacentridae) swimming through coral reef, Post Office Bay, Floreana Island, Galapagos Islands, Ecuador. Pacific Ocean
Flock of Snow geese (Anser caerulescens) leaving their roost at sunrise, Bosque del Apache Wildlife Refuge, New Mexico, USA. January
Aerial view of migrating Wildebeest (Connochaetes taurinus) on the border of Ngorongoro Conservation Area and Serengeti National Park, Tanzania, Africa
American flamingo (Phoenicopterus ruber) flock, feeding in lake at sunset, Rio Lagartas Biosphere Reserve, Yucatan Peninsula, Mexico
Flock of Cobalt-winged parakeets (Brotogeris cyanoptera) feeding on clay at a clay lick, Yasuni National Park, Orellana Province, Ecuador
King penguin (Aptenodytes patagonicus) colony at dawn, Salisbury Plain, South Georgia