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African buffalo (Syncerus caffer) herd showing defensive behaviour as an African lioness (Panthera leo) walks by, Mana Pools National Park, Zimbabwe. September
A school of Siganids (Siganus rivulatus) fish swimming over meadow of little Neptune seagrass (Cymodocea nodosa). Agia Pelagia, Heraklion, Crete, Greece
Horses running in grassland. Bashang Grassland, near Zhangjiakou, Hebei Province, Inner Mongolia, China. July
RF-Horses running through Bashang Grassland. Near Zhangjiakou, Hebei Province, Inner Mongolia, China. July. (This image may be licensed either as rights managed or royalty free)
Banggai Cardinalfish (Pterapogon kauderni), a shoal sheltering against Anchor Coral (Euphyllia ancora). Lembeh Strait, North Sulawesi, Indonesia
RF - Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) waiting to cross the Mara River. Serengeti National Park, Tanzania. September
Nile crocodile (Crocodylus niloticus) waiting to attack White-bearded wildebeest (Connochaetes taurinus albojubatus) as they cross the Mara River
Smallmouth grunt (Haemulon chrysargyreum) fish swim over a coral reef, Cancun, Mexico. Slow shutter speed image
Laughing Gull (Larus atricilla) flock in flight, Sisal, Yucatan Peninsula, Mexico, January
A school of filefifsh take shelter and feed on invertebrates (bottom of frame) found on sargassum in the Sargasso Sea
RF - Impala (Aepyceros melampus) at water with reflections, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
RF - Herd of American Bison (Bison bison) around geo-thermal features. Firehole River Valley. Yellowstone National Park, Wyoming, USA. January 2008
Group of Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Greater flamingo (Phoenicopterus roseus) flock, Pont Du Gau Park, Camargue, France
Springbok (Antidorcas marsupialis) herd on the move, Kgalagadi transfrontier park, South Africa
Alcedo giant tortoise (Chelonoidis vandenburghi), group bathing. Alcedo Volcano, Isabela Island, Galapagos. 2017
Guanacos (Lama guanicoe), five silhouetted on slope in evening. Torres del Paine National Park, Patagonia, Chile. December
Spinner dolphin (Stenella longirostris) pod socialising close to surface. Fury Shoal, Sataya Reef, Marsa Alam, Egypt
Southern royal albatross (Diomedea epomophora) sub-adults, group displaying in tussock grassland. Campbell Island, New Zealand. February
Snow partridge (Lerwa lerwa) flock flying past snowy rockface, in evening light. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana
Caribbean reef shark (Carcharhinus perezi) patrolling coral reef with Schoolmaster snapper (Lutjanus apodus) and other fish. Caribbean Sea off Gardens of the Queen National Park, Cuba
Scarab beetles sleeping overnight in Persian marsh orchid (Dactylorhiza umbrosa). Turkey. June
Red mucronata (Balanophora harlandii), a parasite on woody roots with flowers resembling fungi. Wild Elephant Valley, Xishuangbanna, Yunnan, China
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Ornamental koi carp (Cyprinus carpio) group with open mouths at lake surface. Xishuangbanna Tropical and Flowers Garden, Jinghong, Yunnan, China
Dead leaf / Orange oak leaf (Kallima inachus) butterflies feeding on sap on tree trunk. Mimic leaves with wings closed. Wild Elephant Valley, Xishuangbanna, Yunnan Province, China
African lion, (Panthera leo) confronted by a herd of African buffalo / Cape buffalo (Syncerus caffer), Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Grayling (Thymallus thymallus) in the upper reaches of the Lena River. Baikalo-Lensky Reserve, Siberia, Russia. August 2018
Pod of False killer whales (Pseudorca crassidens) swimming beneath the surface of the ocean. Indian Ocean, off Sri Lanka
Pod of Sperm whale socializing, (Physeter macrocephalus) Dominica, Caribbean Sea, Atlantic Ocean
Fallow deer (Dama dama) herd at dawn, Holkham, Norfolk, England, UK, October
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Emperor penguin (Aptenodytes forsteri) colony, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Brown noddies (Anous stolidus) perched on rock face, Punta Vicente Roca, Isabela Island, Galapagos, Ecuador
Mineral lake with White-cheeked pintails (Anas bahamensis) during quiet volcanic period, on volcanic caldera, Fernandina, Galapagos, Ecuador
Roseate spoonbill (Platalea ajaja) at its nesting colony site, in early morning light. St. Johns Management Area, Florida, USA, April
Red deer (Cervus elaphus) herd of female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) female does and young crossing water, Jura, Scotland, UK, September
Red deer (Cervus elaphus) stag bellowing, surrounded by hinds, Surrey, UK, September
Grey seal (Halichoerus grypus) pup on a breeding beach, Orkney, Scotland, UK, April
Grey seal (Halichoerus grypus) huge group hauled out on a beach, Island of Mingulay, west coast of Scotland. May
Grey seal (Halichoerus grypus) using freshwater pools along the shore line and in the adjacent fields as place to learn to swim, Orkney, Scotland, UK, April
Close up of a colony of Jewel anemones (Corynactis viridis) Scotland, UK, October
Family of six Mute swans (Cygnus olor) at sunrise, silhouetted in waters of Lamlash Bay, Isle of Arran, South Arran Marine Protected Area, Scotland, UK, August
The Bass Rock home to large Northern gannet (Morus bassanus) Firth of Forth, Scotland, UK. August
RF - Mute swans (Cygnus olor) flock of non breeding adults, Fleet lagoon Dorset, England, UK, June
Himalayan ibex (Capra sibirica) during a snowfall, coming down the mountain at dusk to sleep at lower altitude, around Kibber village, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains