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Lemon shark (Negaprion brevirostris) pup and school of fish swimming through Red mangrove (Rhizophora mangle) nursery, Eleuthera Island, Bahamas, North Atlantic
Orbiculate Cardinalfish (Sphaeramia orbicularis) school sheltering amongst roots of mangroves (Rhizophora sp)Orbiculate Cardinalfish (Sphaeramia orbicularis) school sheltering amongst roots of mangroves (Rhizophora sp.) Gam Island, Raja Ampat Islands, West Papua, Indonesia
Three Spotted hyenas (Crocuta crocuta) fighting over meat from a nearby carcass with flock of White-backed vultures (Gyps africanus) watching in background, Mana Pools National Park, Zimbabwe
A breeding herd of African savanna elephants (Loxodonta africana) navigates burning floodplains, Okavango Delta, Botswana
School of orange Scalefin anthias (Pseudanthias squamipinnis) on a coral reef with table corals (Acropora sp)School of orange Scalefin anthias (Pseudanthias squamipinnis) on a coral reef with table corals (Acropora sp.) and red soft corals (Dendronephthya sp.), Ras Mohammed National Park, Sinai, Egypt
Whitetip reef shark (Triaenodon obesus) cruising over a school of Blue and gold snappers (Lutjanus viridis) in a cavern, San Benedicto Island, Revillagigedo Islands, Mexico, Pacific Ocean
Great white egret (Ardea alba) displaying in rookery at Wakodahatchee Wetlands, Florida, USA
Shoal of fish swimming around an outfall pipe on the seabed discharging treated water, Tenerife, Canary Islands, Atlantic Ocean
Large school of juvenile Fusilier (Caesionidae) fish swimming in water column, Andaman Sea, Mu Koh Similan National Park, Phang-nga, Thailand
Shoal of Mediterranean Fairy basslet (Anthias anthias) swimming between Red gorgonian (Paramuricea clavata) that have colonised the crossbars of a sunken electricity pylon that fell into the sea
Flock of White-faced whistling duck (Actophilornis africanus) in flight, Allahein River, The Gambia
School of Bonito fish (Sarda sarda) attacking a school of Spanish sardines (Sardinella aurita) formed into a tight baitball, Isla Mujeres, Mexico, Gulf of Mexico
Atlantic bonito (Sarda sarda) attacking a school of Spanish sardines (Sardinella aurita) which are trapped at the surface, Isla Mujeres, Mexico, Gulf of Mexico
Three Atlantic sailfish (Istiophorus albicans) hunting cooperatively to force a school of Spanish sardine (Sardinella aurita) into a tight baitball and trapping them at the surface, Isla Mujeres
Mute swans (Cygnus olor) in flight against a cloudy sky, Caerlaverock Wetland Centre, Dumfries & Galloway, Scotland, UK. November
RF - Oriental sweetlips fish (Plectorhinchus vittatus) school swimming above a coral reef, Laamu Atoll, Maldives, Indian Ocean. (This image may be licensed either as rights managed or royalty free.)
School of Maldives anemonefish (Amphiprion nigripes) congregating around a Magnificent sea anemone (Heteractis magnifica) on a coral reef, Laamu Atoll, Maldives, Indian Ocean
Powderblue surgeonfish (Acanthurus leucosternon) school swimming over a coral reef, Laamu Atoll, Maldives, Indian Ocean
School of Black rockfish (Sebastes melanops) sheltering in Bull kelp forest (Nereocystis luetkeana) in Browning Pass, British Columbia, Canada. North East Pacific Ocean. September
Scarlet Ibises (Eudocimus ruber) flying onto the top of a mangrove tree in the Orinoco River Delta, Venezuela
Large school of Munks devil rays (Mobula munkiana) aggregating. West Coast of Baja California Peninsula, Mexico. Pacific Ocean
California sea lion (Zalophus californianus) feeding on Sardine bait ball (Sardinops sagax). West Coast of Baja California Peninsula, Mexico. Pacific Ocean
Striped marlin (Tetrapturus audax) feeding on Sardine bait ball (Sardinops sagax). West Coast of Baja California Peninsula, Mexico. Pacific Ocean
Whale shark (Rhincodon typus), an endangered species, feeding on zooplankton in nutrient-rich murky water, being followed by a school of Cobia (Rachycentron canadum), Adang-Rawi Archipelago, Satun
Large school of Bigeye trevally (Caranx sexfasciatus) swimming midwater, Adang-Rawi Archipelago, Andaman Sea, Satun, Thailand
Blade fire coral (Millepora complanata) colonies breaking wave, protection against hurricanes. Guanaja Island, Honduras. Caribbean Sea
Diver inspecting Staghorn coral (Acropora cervicornis) and Finger coral (Porites porites) colonies. Roatan Island, Honduras. Caribbean Sea
Pod of Spinner dolphin (Stenella longirostris) swimming. Utila Island, Honduras. Caribbean Sea
Caribbean flamingo (Phoenicopterus ruber), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Masai giraffe (Giraffa camelopardalis tippelskirchi) herd at sunrise, Masai-Mara Game Reserve, Kenya. November
Juvenile Southern elephant seals (Mirounga leonina) practice fighting near a King penguin (Aptenodytes patagonicus) colony, Gold Harbour, South Georgia Island
A school of Shoulderbar soldierfish (Myripristis kuntee) swimming close to the reef for prtection during the day, Hawaii, Pacific Ocean
Drone aerial view of group of African forest elephants (Loxodonta cyclotis) searching for mineral salts in Dzanga-Sangha National Park, Central African Republic
Flock of Sanderlings (Calidris Alba) in flight with wind turbines in background, Atlantic Coast, Vendee, France. December
Blackbar soldierfish (Myripristis jacobus) sheltering inside small cave, emerging from it at night to forage for food, Scotts Head, Dominica; Eastern Caribbean
A huge school of Pacific cownose / Golden cownose rays (Rhinoptera steindachneri), Baja California, Sea of Cortez, Mexico
Caribbean flamingo (Phoenicopterus ruber) landing, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January
Caribbean flamingo (Phoenicopterus ruber) flock, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January
Small flock of migrating Black-tailed godwits (Limosa limosa) in flight against a bright blue sky, River Coquet Estuary, Northumberland, UK. September
Red knot (Calidris canutus) flock at high tide roost. Snettisham, Norfolk, UK. October
Common starling (Sturnus vulgaris) murmuration in between high voltage powerlines and electricity pylons, The Netherlands, Europe. February
School of Yellow-ribbon sweetlips (Plectorhinchus polytaenia) swimming with the current, Raja Ampat, Indonesia, Pacific Ocean
Spinifex pigeon (Geophaps plumifera) flock on stony desert floor, Diamantina River, Queensland, Australia
Galah (Eolophus roseicapillus) flock of birds flying over grassland, Diamantina River, Queensland, Australia
Eider ducks (Somateria mollissima) floating on waves, Iceland. March
Snow bunting (Plectrophenax nivalis) flock resting on a roof, Iceland. March
Iceland horses in winter, western Iceland. March
Aerial view of Caribbean flamingo (Phoenicopterus ruber) breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate