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Himalayan ibex (Capra sibirica) during a snowfall, coming down the mountain at dusk to sleep at lower altitude, around Kibber village, Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains
Shoal of Piraputanga, (Brycon hilarii), close to the surface, Aquario Natural, Bonito, Mato Grosso do Sul, Brazil
Shadowfin soldierfish (Myripristis adusta) West Papua, Indonesia
Shrimpfish / Razorfish (Aeoliscus punctatus / strigatus) have adapted to a vertical posture and swim with their head downward. Lembeh, North Sulawesi, Indonesia
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Red-and-green macaws (Ara chloropterus) perched and in flight over Buraco das Araras, the Sinkhole of the Macaws, Jardim, Mato Grosso do Sul, Brazil. September
Aerial view of African elephants (Loxodonta africana) looking for the shade under trees on a very hot day, Northern Botswana
Plain / Burchells zebras in dust (Equus burchelli) Nxai Pan National Park, Botswana
Brandts cormorant (Phalacrocorax penicillatus) hunts for a meal in a school of Pacific chub mackerel (Scomber japonicus), beneath an oil rig
King penguin (Aptenodytes patagonicus) chick creche, Volunteer Point, East Falkland, Falkland Islands, October
King penguin (Aptenodytes patagonicus) colony, Volunteer Point, East Falkland, Falkland Islands, October
Zebra at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Whooper swans (Cygnus cygnus), flying at sunset, Caerlaverock Wildfowl & Wetland Trust, Dumfries & Galloway, Scotland, November 2016
Shadowfin soldierfish (Myripristis adusta) schooling over coral reef. Dampier Strait, Raja Ampat, West Papua, Indonesia, March 2016
South American sea lion (OtariaAflavescens) colony on beach, Punta Norte, Peninsula ValdesSouth American sea lion (Otariaflavescens) colony on beach, Punta Norte, Peninsula Valdes, Argentina
Northern gannets (Morus bassanus). Bass Rock, Scotland, UK. August
Mute swan (Cygnus olor) flock on snow, Hazerswoude, The Netherlands, February
Atlantic Puffins (Fratercula arctica) in flight in mist, Skomer Island, Wales, UK, August
Blue wildebeest (Connochaetes taurinus), migrating herd silhouetted on the horizon at sunset, Masai Mara Game Reserve, Kenya
Mobula / Chilean devil ray (Mobula tarapacana) with a school of fish (Capros aper) showing vertical migration, Santa Maria, Azores, Portugal, Atlantic Ocean
Soft corals (Dendronephthya hemprichi) growing in very shallow water in the shade provided by a jetty, while a school of Silversides (Atherinomorus lacunosus) circle. Berenice Jetty, Aqaba, Jordan
Silversides (Atherinomorus lacunosus) school just below surface in evening light, Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Gannet (Morus bassanus) nearest out of focus, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Gorgonian sea fans (Melithaea sp. ) with a large school of Pygmy sweepers (ParapriacanthusGorgonian sea fans (Melithaea sp.) with a large school of Pygmy sweepers (Parapriacanthus ransonetti) Andaman Sea, Thailand
Lesser flamingos (Phoeniconaias minor) flock on lake at dawn, Lake Ndutu Tanzania
Red snappers / Two-spot red snappers (Lutjanus bohar) mass shoal of fish spawning, Palau, Philippine Sea. A few black and white snappers (Macolor niger) have crawled into the school to eat the eggs
Crevalle Jack (Caranx hippos) and other fish entering Homosassa Spring for the winter, Homosassa Springs Wildlife Park, Florida, USA. March. Photographed for the Freshwater Project
European bison / Wisent (Bison bonasus) herd released into the Tarcu mountains nature reserve, Natura 2000 area, Southern Carpathians, Romania. May 2014
RF- Caribou herd (Rangifer tarandus) grazing on tundra. Kobuk Valley National Park, Alaska, USA
School of Bonito (Sarda orientalis) in open water. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Bigeye trevally (Caranx sexfasciatus) forming a circular shoal in blue water, on the drop off of a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Whitebar surgeonfish (Acanthurus leucopareius) school in undewater cave, oxygenated water brought down by Pacific ocean surge. Kona, The Big Island, Hawaii
Long Beaked Common Dolphin (Delphinus capensis), Espiritu Santo National Park, Sea of Cortez (Gulf of California), Mexico, February
Griffon vulture (Gyps fulvus) landing amongst other vultures on cliff above sea, Liendo Valley within the Montana Oriental Costera, Cantabria, Spain
Lyretail anthias / Goldies (Pseudanthias squamipinnis) on reef top. Egypt, Red Sea
Bearded tit (Panurus biarmicus), five perched on Reed (Phragmites australis). Danube Delta, Romania. May
Southern elephant seal (Mirounga leonina), male at sunrise with King penguin (Aptenodytes patagonicus) colony in background. St Andrews Bay, South Georgia. October 2017
Family of Whooper swan (Cygnus cygnus), Iceland, February
Razorbill (Alca torda), pair on cliff, Farne Islands, Northumberland, England, UK, June
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December
School of Scaled herring or Scaled sardine (Harengula clupeola), entering from the sea into the freshwater of the Laguna Yal Ku, Quintana Roo, Yucatan Peninsula, Mexico
Chub (Squalius cephalus), on spawning ground Sava Bohinjka River, Slovenia June 2013. Photographed for The Freshwater Project
Lesser flamingo (Phoeniconaias minor) flock, Walvis Bay Namibia
Wild yak herd (Bos mutus) Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
Peach-fronted parakeets (Aratinga aurea) at the nesthole in termite mount, Pantanal, Brazil
