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Mallard (Anas platyrhynchos) male flying low over water. Cambridgeshire Fens, England, March
Hen harrier (Circus cyaneus) female hovering over moorland, Glen Tanar Estate, Deeside, Scotland, UK, June
Stellers sea-eagle (Haliaeetus pelagicus) landing on pack ice, Hokkaido, Japan, February
Barn owl (Tyto alba) in flight, captive, Cumbria, UK
Booted Racket-tail hummingbird (Ocreatus underwoodii) male flying, feeding from flower, Mindo, Ecuador, Andes, South America, January
Arctic tern (Sterna paradisaea) hovering above the ocean, Svalbard, Norway June
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Little Owl (Athene noctua) taking flight from a post. Wales, UK, June
Oystercatchers (Haematopus ostralegus) arriving to roost at Snettisham on the Wash. Norfolk, January
Atlantic Puffins (Fratercula arctica) flying near cliff top, Isle of Lunga, Isle of Mull, Treshnish Isles, Scotland, UK, June
Black-headed gull (Chroicocephalus ridibundus) in flight, artistically blurred photograph, Cheshire, UK, February. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Atlantic puffin (Fratercula arctica) gathering around burrows on cliff top, Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland, UK, June
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009
White-tailed eagle (Haliaeetus albicilla) taking off with fish prey, Norway, August
Eurasian crag martin (Ptyonoprogne rupestris) Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May
White chinned petrel (Procellaria aequinoctialis) in flight low over the water off Kaikoura, Canterbury, New Zealand
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Small Tortoiseshell Butterfly (Aglais urticae) feeding on scabious flowers in hay meadow, Wensum Valley, Norfolk, England, UK, July
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
Lapwings (Vanellus vanellus), Golden plover (Pluvialis apricaria), Wigeon (Anas penelope) and Common Teal (Anas crecca) flying over flooded marshes in winter, with a rainbow in the background
Flock of Manx shearwaters (Puffinus puffinus) in flight over sea, near Ardnamurchan Point, Inner Hebrides, Scotland, UK, July 2011
Hen harrier (Circus cyaneus) adult female landing on rock in moorland habitat, Glen Tanar Estate, Grampian, Scotland, UK, June
Hen Harrier (Circus cyaneus) adult female in flight over heather moorland, Scotland, UK. July
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Kingfisher (Alcedo atthis) taking off from water with caught fish. Worcestershire, UK, March
Common starling (Sturnus vulgaris) flocking prior to roosting, Scotland, January 2009
Golden eagle (Aquila chrysaetus) Kuusamo, Finland, January
King Eider (Somateria spectabilis) flying low over water. Floe edge, Arctic Bay, Nunavut, Canada, April
Golden eagle (Aquila chrysaetos) adult female taking off, flying over mountain landscape, trained bird photographed during filming in Glenveagh National Park, Donegal, Ireland. August 2010
Broad-tailed Hummingbird {Selasphorus platycercus} male in flight feeding on Musk Thistle (Carduus nutans) Rocky Mountain National Park, Colorado, USA, June
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Common buzzard (Buteo buteo) in flight with hoar frost. Northamptonshire, UK, January
Lammergeier / Bearded vulture (Gypaetus barbatus) in flight with the out-of-focus moon behind. Ladakh, Himalayas, northern India
RF- Carrion crow (Corvus corone) flying from a snow covered pine tree in a winter landscape. Valsavarenche, Gran Paradiso NP, Italy, March
Griffon vulture (Gyps fulvus) in flight, Extremadura, Spain, April 2009
Violet sabrewing hummingbird (Campylopterus hemileucurus) male hummingbirds territorial fight at dawn, Turrialba, Costa Rica
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Helicopter flying over volcanic eruption near Eyjafjallajoekull glacier, Iceland, 24th March 2010. Volcano previously dormant since 1821
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Sword-billed hummingbird (Ensifera ensifera) hovering in flight, North-Ecuador
Wood pigeon (Columba palumbus) flying with nesting material in beak, Lorraine, France, April
Great grey owl (Strix nebulosa) hunting over snow, woodland in background. Kuusamo, Finland. February
