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Avocet (Recurvirostra avosetta) trying to drive Shelduck (Tadorna tadorna) pair away from breeding grounds, Oare marshes, RSPB Greater Thames Futurescapes Project, North Kent, UK, June
Chilean flamingos (Phoenicopterus chilensis) group of three in flight, Lago Amarga, Torres del Paine National Park, Patagonia, Chile
Black grouse (Tetrao tetrix) male in flight, Cairngorms National Park, Scotland, UK, April
Barn swallow (Hirundo rustica) alighting at pond to collect mud for nest building, Scotland, UK, May
Peregrine falcon (Falco peregrinus), adult male in flight. Bristol, UK. March
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
Rufous-tailed hummingbird (Amazilia tzacatl) territorial fighting around Heliconia flower (Heliconia latispatha) La Selva, Costa Rica
Tree swallow (Tachycineta bicolor) in fight over the Madison River, Montana, USA, June
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Garden bumblebee (Bombus hortorum), flying to Foxglove (Digitalis purpurea), flower, Monmouthshire, Wales, UK. June
Honeybee (Apis mellifera) flying to Bluebell flowers (Hyacinthoides non-scripta) Monmouthshire, Wales, UK. April
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
European honey bee (Apis mellifera) in flight, feeding on flowers (Brachyglottis sp). Monmouthshire, Wales, UK, June
White-tailed eagle (Haliaeetus albicilla) in flight, Norway, April
Canada geese (Branta canadensis) flock in flight, Moray Firth, Highlands, Scotland. June
RF -Three colourful Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
RF - Four colourful Red-and-green macaws or Green-winged macaws (Ara chloropterus) in flight over forest canopy. Mato Grosso do Sul, Brazil. September
Red-and-green macaw (Ara chloropterus) in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
Great Grey Owl (Strix nebulosa) landing in snow, Kuusamo, Finland
Long eared owl (Asio otus) in flight, taking off from oak tree snag. Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany
Snowy owl (Bubo scandiacus) in flight, Boundary Bay, British Columbia, Canada. February. First place in the Overall Category of the 2013 Great Backyard Bird Count photo competition
Great Grey owl (Strix nebulosa) flying through woodlands, Tornio, Finland, Scandinavia, March
Great Grey Owl (Strix nebulosa) landing, Hyvink Finland, January 2007
Long eared owl (Asio otus) in flight with snow. Germany, Europe
African fish eagle (Haliaeetus vocifer) flying, Selinda Spillway, northern Botswana
Imperial Eagle (Aquila heliaca) juvenile flying low, Hungary January
Steppe eagle (Aquila nipalensis) in flight, Oman, November
Golden Eagle (Aquila chrysaetos) in flight, snowing, Utajarvi, Finland, February
Golden Eagle (Aquila chrysaetos) coming in to land with claws spread. Kuusamo, Finland, February
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Flatanger, Norway, June
Golden eagle (Aquila chrysaetos) taking off, Flatanger, Norway, November 2008
RF- Golden eagle (Aquila chrysaetos) landing in snow, Flatanger, Norway. November
Golden Eagle (Aquila chrysaetos) in flight. Utajrvi, Finland. February
Brown long eared bat (Plecotus auritus) in flight, Surrey, UK
Common tern (Sterna hirundo) in flight over water, Hampstead Heath, London, UK, August
Two Goldfinches (Carduelis carduelis) squabbling over Common teasel (Dipsacus fullonum) seeds in winter, Hope Farm RSPB reserve, Cambridgeshire, England, UK, February. 2020VISION Exhibition
Kingfisher (Alcedo atthis) rising from water after diving for prey. Worcestershire, UK, March
Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January
Northern gannet (Morus bassanus) colony in mist, Hermaness, Shetland Isles, Scotland, July 2009
Golden eagle (Aquila chrysaetos) sub-adult male (two years) flying down to take prey, Cairngorms National Park, Scotland, UK, captive
Greater horseshoe bat {Rhinolophus ferrumequinum} in flight hunting a moth at night, Germany
RF - Marbled white butterfly (Melanargia galathea) on knapweed, with soft focus leaves, Aosta Valley, Gran Paradiso National Park, Italy
Glorious scarab (Chrysina gloriosa) flying, Brewster County, Texas, USA. Controlled conditions. August
Lesser flamingo flock (Phoeniconaias) flying over lake with green algae, aerial view, Rift valley, Kenya
Black-headed gull (Chroicocephalus ridibundus) alighting on water at dawn, Cheshire, UK, November. Highly Commended in the British Wildlife Photography Awards (BWPA) competition 2013
Buff-tailed bumblebee (Bombus terrestris) flying to Oriental poppy (Papaver orientale) Monmouthshire, Wales, UK. July
Whale headed stork (Balaeniceps rex) taking off, Lake Albert, Uganda, East Africa
Caribbean flamingo (Phoenicopterus ruber) about to take off to roosting site, Ria Celestun Biosphere Reserve, Yucatan Peninsula Mexico, January