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White stork (Ciconia ciconia) pair before mating at nest site on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
Gannet (Morus bassanus) courtship, Saltee Islands, County Wexford, Ireland, June 2009
Gannets (Morus bassanus) Saltee Islands, County Wexford, Ireland, June 2009
Grey seal (Halichoerus grypus) mating behaviour, Donna Nook, Lincolnshire, UK, November 2008
Skylark (Alauda arvensis) singing on branch stump, East Slovakia, Europe, June 2008
European Roller (Coracias garrulus) pair, display, Pusztaszer, Hungary, May 2008
Turtle Dove (Streptopelia turtur) pair, Pusztaszer, Hungary, May 2008
Red-backed shrike (Lanius collurio) pair, male feeding insect to female in courtship, Bulgaria, May 2008
Common roller (Coracias garrulus) female feeding on its courtship gift, Bulgaria, May 2008
European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 1 of 2
Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male in flight at lek, Cairngorms NP, Grampian, Scotland, UK, April
Pair of Gannets (Morus bassanus) mutual preening, Bass Rock, Firth of Forth, Scotland, UK, June
Black grouse (Tetrao tetrix) male displaying at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
Black necked grebe (Podiceps nigricollis) pair at night, performing their courtship dance during the mating season, with street lights reflected in the water. The Netherlands.April 2014
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Brown hare (Lepus europaeus) two males chasing female in snow covered field during a blizzard. Derbyshire, England, UK, February. Winner of British Seasons category of BWPA competition 2014
Moorland Clouded Yellow butterfly (Colias palaeno) male and female, Joutsa (formerly Leivonmaki), Finland, July
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Great crested grebe (Podiceps cristatus) pair performing courtship display at dawn, backlit with surrounding by mist, Cheshire, UK, March
RF- Black Grouse (Lyrurus tetrix) displaying. Finland. March
Common cranes (Grus grus) displaying, Hula Valley, Northern Israel, January
Silhouette of two male Black grouse (Tetrao tetrix) displaying at lek at dawn, Cairngorms NP, Grampian, Scotland, UK, April. 2020VISION Book Plate
Black Grouse (Tetrao tetrix) males at spring lek. Deeside, Scotland, April
Male Black grouse (Tetrao tetrix) displaying at lek at dawn, Cairngorms NP, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male at lek, Creag Megaidh NNR, Highlands, Scotland, UK, April
Capercaillie (Tetrao urogallus) male displaying in pine forest in snow, Cairngorms NP, Highlands, Scotland, UK, January. Photographer quote
Indigo hamlets (Hypoplectrus indigo) pair spawning at dusk, George Town, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea
Capercaillie (Tetrao urogallus) male, displaying in pine forest, Cairngorms NP, Scotland, UK, April
Andean cock of the rock (Rupicola peruviana) male in courtship ritual, Andean Mountains, Merida, Venezuela
Wolf Spiders (Pardosa sp. ), male RIGHT waving palps in courtship displayWolf Spiders (Pardosa sp.), male RIGHT waving palps in courtship display. Peak District National Park, Derbyhsire, UK. May
Northern gannets (Morus / Sula bassanus) portrait of individual with a courting pair in foreground, UK, Bass Rock, Scotland
Male Indian peafowl (Pavo cristatus) displaying, captive
Couchs Spadefoot (Scaphiopus couchii) adult at night calling with vocal sac inflated. Laredo, Webb County, South Texas, USA, April
A pair of Ravens (Corvus corax) perched in bleached out pine tree, Estonia, March 2008
Pair of King Penguins (Aptenodytes patagonicus) on Gold Beach, South Georgia Island, Sub Antarctica
Great crested grebes (Podiceps cristatus) performing their elaborate courtship ritual at dawn, Cheshire, UK November
