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Leopard (Panthera pardus) pair mating behaviour, Okavango Delta, Botswana
Black Grouse (Lyrurus tetrix) fighting on icem Finland. March
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Eurasian rollers (Coracias garrulus) courting on branch, Hungary. June
King cormorants (Phalacrocorax atriceps) in courtship behaviour at their nest site. Saunders Island, Falkland Islands
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Common cranes displaying by woodland {Grus grus} Finland
Fallow deer (Dama dama) herd at dawn, Holkham, Norfolk, England, UK, October
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Black-browed albatross pair (Thalassarche melanophris) mutual grooming during copulation. New Island, Falkland Islands, November
Nurse sharks (Ginglymostoma cirratum) three in a courtship dance at sunrise in a mangrove area near Eleuthera, Bahamas
Magellanic penguin (Spheniscus magellanicus) calling at the entrance of burrow, courtship display, Carcass Island, Falkland Islands, October
Laysan albatross (Phoebastria immutabilis), courtship ritual, Eastern island, Midway Atoll National Wildlife Refuge, Hawaii
Brown Hare (Lepus europaeus), Bayern, Germany. April
Green mandarinfish (Synchiropus splendidus) pair courting and mating at sunset, Sulu Sea, Philippines
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Anemone Hermit Crab (Pagurus prideaux) courtship behaviour. Maseline Harbour, Sark, British Channel Islands, September
Northern gannets {Morus bassanus} pair in courtship display, Great Saltee, Co. Wexford, Ireland, June
Capercaillie (Tetrao urogallus) male displaying and calling at lek, Kamchatka, Far east Russia, May
Long-tailed manakin (Chiroxiphia linearis) perched on lek (display perch), Costa Rica
Sooty Albatross (Phoebetria fusca) group courting on high inland ridge. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic. Endangered
RF - Red deer (Cervus elaphus) stags fighting, Holkham Park, Norfolk, England, UK, September
Black Grouse (Tetrao tetrix) lek with male displaying and females around in winter, Tver, Russia. April
Black Grouse (Tetrao tetrix) males fighting in winter, Tver, Russia. April
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Mediterranean Monk seal (Monachus monachus) pair courting on rocks on seabed, Deserta Grande, Desertas Islands, Madeira, Portugal, August 2009, Critically endangered species
White stork (Ciconia ciconia) pair at nest site on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
White stork (Ciconia ciconia) pair, courtship, at nest on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
White stork (Ciconia ciconia) pair at nest engaged in courtship display, male with nesting material, Lithuania, May 2009
Male Spanish ibex (Capra pyrenaica) scenting air behind female with tongue, Sierra de Gredos, Spain, November 2008. WWE OUTDOOR EXHIBITION. NOT AVAILABLE FOR GREETING CARDS OR CALENDARS
Pair of Common hermit crabs (Pagurus bernhardus) male carrying female until she leaves her shell and they can mate, Trondheimsfjorden, Norway, February 2009
White stork (Ciconia ciconia) pair before mating at nest site on old chimney, Rusne, Nemunas Regional Park, Lithuania, June 2009
Gannet (Morus bassanus) courtship, Saltee Islands, County Wexford, Ireland, June 2009
Gannets (Morus bassanus) Saltee Islands, County Wexford, Ireland, June 2009
Grey seal (Halichoerus grypus) mating behaviour, Donna Nook, Lincolnshire, UK, November 2008
Skylark (Alauda arvensis) singing on branch stump, East Slovakia, Europe, June 2008
European Roller (Coracias garrulus) pair, display, Pusztaszer, Hungary, May 2008
Turtle Dove (Streptopelia turtur) pair, Pusztaszer, Hungary, May 2008
Red-backed shrike (Lanius collurio) pair, male feeding insect to female in courtship, Bulgaria, May 2008
Common roller (Coracias garrulus) female feeding on its courtship gift, Bulgaria, May 2008
European Hares (Lepus europaeus) boxing, female on right. Wales, UK, June. Sequence 1 of 2
Black grouse (Tetrao tetrix) males fighting at lek site, Cairngorms National Park, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male in flight at lek, Cairngorms NP, Grampian, Scotland, UK, April
