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Common goldeneye (Bucephala clangula) male performing its mating display. Southern Norway. March
Raggiana bird of paradise (Paradisaea raggiana) male displaying to female, Papua New Guinea, Varirata NP
Spotted owlet (Athene brama) pair on a branch, Bharatpur / Keoladeo Ghana National Park, India
Red deer (Cervus elaphus) stag bellowing during rut, at night with full moon, Derbyshire, UK
Kingfisher (Alcedo atthis) male passing fish to female spring courtship behaviour, Hertfordshire, England, UK, March. Sequence 4 of 6
Guianan Cock-of-the-rock (Rupicola rupicola), during courtship display, Guianan Shield, Suriname
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Male Carolas Parotia (Parotia carolae) bird of paradise displaying on his display court to five females perched above, Crater Mountain Wildlife Management Area, Eastern Highlands Province
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Mute swan (Cygnus olor) pair courting. Walthamstow reservoir, London, UK
Humpback whale (Megaptera novaeangliae australis) courtship, with the female above showing her abdomen and the male below. Vava'u, Tonga. Pacific Ocean
Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Northern gannet (Morus bassanus) pair, courtship, Heligoland, Germany
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA. When displaying
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
TemminckAśs tragopan (Tragopan temminckii) male during courtship displayTemminck's tragopan (Tragopan temminckii) male during courtship display, showing red-blue lappets. Tibet, China
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October
Manchurian crane (Grus japonensis) pair in courtship dance during snowstorm. Hokkaido, Japan. March
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Eared grebes (Podiceps nigricollis) stand erect in water during courtship, North Park, Colorado, USA. June
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
Fallow deer (Dama dama) buck bellowing in autumn vegetation, Bradgate Park, Leicestershire, UK, October
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Western Grebe (Aechmorphus occidentalis), pair during rushing courtship display in which they run across waters surface in synchrony, near Escondido, California, USA, January
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
Wild Turkeys (Meleagris gallopavo), two toms courting several hens, New York, USA
White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Two rainbow lorikeets (Trichoglossus moluccanus) on branch, Werribee, Victoria, Australia
Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Hoopoe (Upupa epops) male feeding female in courtship display, Pusztaszer, Hungary, April