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Sperm whale (Physeter macrocephalus) young male trying to copulate with a female. The calf of the female was in the way so the male is trying to chase it away. Dominica
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India
Flightless cormorant (Phalacrocorax harrisi) pair in courtship Puerto Pajas, Isabela Island, Galapagos
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Western capercaillie (Tetrao urogallus) male calling at lek, with female, Tver, Russia. May
Black Grouse (Tetrao tetrix) males fighting at lek in the snow, Tver, Russia. April
Tennessee Warbler (Vermivora peregrina) male perched in a pine tree branch, singing in spring, Alberta, Canada. May
Pair of Horned Grebes (Podiceps auritus) in breeding plumage, on water, Southeast Alberta, Canada. May
Gunnison sage-grouse (Centrocercus minimus) male displaying at a lek. Gunnison County, Colorado. April
Royal terns (Sterna maxima) pair crossing bills during courtship behaviour, Fort DeSoto Park, Florida, USA
Hoopoe (Upupa epops) male feeding female in courtship display, La Serena, Extremadura, Spain, April 2009
European bee-eater (Merops apiaster) pair in courtship display, Bulgaria, May 2008
European Hare (Lepus europaeus) courting pair in grassland at dawn. Wales, UK, July
Three European Hare (Lepus europaeus) chasing, a courtship behaviour. Wales, UK, June
Capercaillie (Tetrao urogallus) male displaying. Cairngorms National Park, Scotland, UK, September
Great crested grebe (Podiceps cristatus) pair of adults during part of their elaborate courtship ritual during the very last few rays of evening sunlight, Derbyshire, UK, March
Black grouse (Tetrao tetrix) two males displaying and fighting at lek, Cairngorms NP, Grampian, Scotland, UK, April
Rear view of Black grouse (Tetrao tetrix) male displaying at lek, Cairngorms National Park, Grampian, Scotland, UK, April
Black grouse (Tetrao tetrix) male displaying at lek, Cairngorms National Park, Grampian, Scotland, UK, April
RF- Capercaillie (Tetrao urogallus) male displaying, Inshriach, Cairngorms National Park, Scotland, UK. February
Whooper swan (Cygnus cygnus) pair on loch in heart shape, Cairngorms National Park, Scotland, UK, November
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Satellite male Ruff (Philomachus pugnax) in full display at the lek. Varanger, Finmark, Norway, May
Lesser kestrel (Falco naumanni) male flying past during courtship with Mole cricket in its talons. Extremadura, Spain
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Two Cheetahs (Acinonyx jubatus) touching noses in greeting display, Serengeti NP, Tanzania, East Africa, January
Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Goldies Bird of Paradise (Paradisaea decora) male displaying to female at lek, Fergusson Island, Milne Bay Province, Papua New Guinea. Near-threatened
Mating pair of King Penguins (Aptenodytes patagonicus) on Gold Beach, South Georgia Island, Sub Antarctica
Gannet (Morus bassanus) breeding pair during part of their elaborate courtship ritual. Saltee Islands, Republic of Ireland, May
Waved albatross (Phoebastria irrorata) courting pair mutually grooming. Punta Cevallos, Espanola (Hood) Island, Galapagos, Ecuador, May
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May
Dusky dolphins (Lagenorhynchus obscurus) courting pair leaping from the sea, Kaikoura, Canterbury, New Zealand, January
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Northern gannet (Morus bassanus) pair, courtship, Heligoland, Germany
Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October
Pair of Waved albatrosses (Phoebastria irrorata) billing as greeting. Punta Suarez, Espanola Island, Galapagos Islands, Ecuador
Wine throated hummingbird (Selasphorus ellioti) male, courtship display, Cloud forest, Atitlan Volcano, Solola, Guatemala
Ruddy duck (Oxyura jamaicensis) male in courtship display. Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Humpback whales (Megaptera novaeangliae) pair engaging in courtship behavior, dancing for hours, off Tubuai, French Polynesia, Pacific Ocean
Bluethroat (Luscinia svecica cyanecula) male, singing as a courtship display, Germany
Magnificent frigatebirds (Fregata magnificens) pair in courtship display, male with gular sac inflated, North Seymour Island, Galapagos National Park, Galapagos Islands
