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Little owl {Athene noctua) pair perched, courtship behaviour, Spain
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Rainbow lorikeets (Trichoglossus moluccanus) pair preening each other, perched in tree. Ulladulla, New South Wales, Australia
Vogelkop superb bird-of-paradise (Lophorina niedda) male, performing courtship display on his display log, flashing false blue eye spots, a bright yellow gape, and blue breast shield
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
Male Sage Grouse {Centrocercus urphasianus} courtship display, Baggs, Wyoming, USA
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
European Bee-eater (Merops apiaster) pair, male passing Dragonfly prey to female, Pusztaszer, Hungary, May 2008
Great crested grebe (Podiceps cristatus) pair performing the weed dance, a spectacular part of their elaborate courtship ritual, Derbyshire, UK, March
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Common grackle pair (Quiscalus quiscula) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Wood pigeons (Columba palumbus) male and female during courtship preening display, Norfolk, England, UK, August
Mandarin duck drake (Aix galericulata) on the water, showing orange sail feathers which are used when displaying, Southwest London, UK, March
Clarks grebes (Aechmophorus clarkii) courting pair performing the Weed Ceremony in which they slowly circle breast to breast holding pond weed as nesting material, Lake Hodges, Escondido, California
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia
Common Snipe (Gallinago gallinago) autumn courtship display, Germany, August
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Satin bowerbird male at bower decorated with blue objects to attract mate, Lamington NP
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
Capercaillie (Tetrao urogallus) adult male displaying in pine forest. Cairngorms National Park, Scotland, February
Capercailie (Tetrao urogallus) male displaying from branch of Scots pine tree. Cairngorms National Park, Highlands, Scotland, UK, March
Common goldeneye (Bucephala clangula) male performing its mating display. Southern Norway. March
Raggiana bird of paradise (Paradisaea raggiana) male displaying to female, Papua New Guinea, Varirata NP
