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Little owl {Athene noctua) pair perched, courtship behaviour, Spain
Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Vogelkop superb bird-of-paradise (Lophorina niedda) male, performing courtship display on his display log, flashing false blue eye spots, a bright yellow gape, and blue breast shield
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
European Bee-eater (Merops apiaster) pair, male passing Dragonfly prey to female, Pusztaszer, Hungary, May 2008
Male Sage Grouse {Centrocercus urphasianus} courtship display, Baggs, Wyoming, USA
Mating pair of King Penguins (Aptenodytes patagonicus) on Gold Beach, South Georgia Island, Sub Antarctica
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Great crested grebe (Podiceps cristatus) pair performing the weed dance, a spectacular part of their elaborate courtship ritual, Derbyshire, UK, March
Silvery grebe (Podiceps occipitalis) attempting to join Hooded grebes (Podiceps gallardoi) during a breeding display, Patagonia National Park, Santa Cruz, Argentina. Cropped
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Rainbow lorikeets (Trichoglossus moluccanus) pair preening each other, perched in tree. Ulladulla, New South Wales, Australia
Bower of Vogelkop Bowerbird (Amblyornis inornata) with bright decorations. Nov 2004
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Elk (Cervus canadensis) bull and cw sniffing noses, Yellowstone National Park, Wyoming, USA. September
Rocky mountain elk (Cervus elaphus nelsoni) bull bugling during rut. Rocky Mountain National Park, Colorado, USA. October
Red Bird-of-Paradise (Paradisaea rubra) male performing practice display at tree-top lek, Raja Ampat, Indonesia. *Medium repro only
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Common grackle pair (Quiscalus quiscula) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Sarus crane (Grus antigone), male displaying, Keoladeo NP, Bharatpur, India
Wood pigeons (Columba palumbus) male and female during courtship preening display, Norfolk, England, UK, August
Mandarin duck drake (Aix galericulata) on the water, showing orange sail feathers which are used when displaying, Southwest London, UK, March
Clarks grebes (Aechmophorus clarkii) courting pair performing the Weed Ceremony in which they slowly circle breast to breast holding pond weed as nesting material, Lake Hodges, Escondido, California
Black terns (Chlidonias niger), pair on a perch, vocalizing during courtship (termed Erect Posture display), Montezuma National Wildlife Refuge, New York, USA, May
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia
Common Snipe (Gallinago gallinago) autumn courtship display, Germany, August
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Satin bowerbird male at bower decorated with blue objects to attract mate, Lamington NP
King penguins (Aptenodytes patagonicus) at sunrise. Two males and a female, with the males fighting for the attention of the female. Falklands Islands
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Magnificent bird of paradise (Cicinnurus magnificus) male, leaning back and presenting his breast shield and raised cape to two females, courtship display, Arfak Mountains, West Papua, Indonesia
Wilson's bird of paradise (Cicinnurus respublica) male, courtship display, Batanta Island, West Papua, Indonesia
Pair of Wilson's birds of paradise (Cicinnurus respublica), male performing courtship display to female from a small sapling in the middle of his display court, Batanta Island, West Papua
Pair of Great crested grebes (Podiceps cristatus) performing the weed dance during courtship, The Netherlands, Europe. March
Pair of Waved albatross (Phoebastria irrorata) courtship, Espanola Island, Galapagos, South America
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Greater Prairie-Chicken (Tympanuchus cupido) male displaying on lek. Fort Pierre National Grassland, South Dakota, USA
Red fox (Vulpes vulpes) pair nuzzling. London, UK. February
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Boat-tailed grackle pair (Quiscalus major) in courtship display. Wakodahatchee Wetlands, Florida, USA, April
Lawess parotia (Parotia lawesii) male displaying. Papua New Guinea
TemminckAśs tragopan (Tragopan temminckii) male during courtship displayTemminck's tragopan (Tragopan temminckii) male during courtship display, showing red-blue lappets. Tibet, China
Red Deer (Cervus elaphus) stags fighting during the rutting season. Bushy Park, London, UK. October
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Pair of courting Egg-eating sea snakes / Turtleheaded sea snakes (Emydocephalus annulatus) New Caledonia, Pacific Ocean
Manchurian crane (Grus japonensis) pair in courtship dance during snowstorm. Hokkaido, Japan. March
Eared grebes (Podiceps nigricollis) stand erect in water during courtship, North Park, Colorado, USA. June
Amur / Siberian tiger (Panthera tigris altaica) pair in courtship, in snow. Captive in tiger park, Heilongjiang Province, China. January
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
Fallow deer (Dama dama) buck bellowing in autumn vegetation, Bradgate Park, Leicestershire, UK, October
Waved albatross (Phoebastria irrorata) pair in courtship display at nest site, Punta Suarez, Espanola Island, Galapagos
Great frigatebird (Fregata minor) male displaying inflated gular sac, Galapagos, Ecuador
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Lioness (Panthera leo), aggressively rebuffing male lion banks of the Luangwa river, South Luangwa National Park, Zambia
Northern gannets (Morus bassanus) performing sky-pointing display with outspread wings as part of courtship, Cape St. Marys Ecological Reserve, Newfoundland, Canada. June
Red-breasted Mergansers (Mergus serrator) male in foreground performing courtship display to female in background (left), Aurora, New York, USA, April
Western Grebe (Aechmorphus occidentalis), pair during rushing courtship display in which they run across waters surface in synchrony, near Escondido, California, USA, January
American oystercatchers (Haematopus palliatus) courting pair performing Piping Display, calling as they walk along together, Fort DeSoto Park, Florida, USA, March
Wild Turkeys (Meleagris gallopavo), two toms courting several hens, New York, USA
White stork (Ciconia ciconia) pair performing an up-down display with bill clattering on their nest in an Oak tree, Knepp estate, Sussex, UK, April 2019
Red deer (Cervus elaphus) herd during rut in morning light. Derbyshire, England, UK. October
Jaguars (Panthera onca) courting pair on a sand bank. Cuiaba River, Northern Pantanal, Mato Grosso, Brazil
Two rainbow lorikeets (Trichoglossus moluccanus) on branch, Werribee, Victoria, Australia