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Red squirrel (Sciurus vulgaris) feeding in snow. Scotland, UK. March
Red squirrel (Sciurus vulgaris) jumping towards camera. Scotland, UK. February
Marsh fritillary (Euphydryas aurinia), group feeding on Marsh thistle (Cirsium palustre). Lincolnshire, England, UK
Rivolis hummingbird (Eugenes fulgens) nectaring on Fuchsia (Fuchsia sp) flower. Costa Rica
Green-crowned brilliant (Heliodoxa jacula), three feeding on Lobster-claw (Heliconia sp) flower in rain. Costa Rica
Red fox (Vulpes vulpes) cub looking over shoulder at camera. Sheffield, England, UK. May
Red fox (Vulpes vulpes) cub looking at camera, in morning. Sheffield, England, UK. June
Turtle dove (Streptopelia turtur) standing at a drinking pool. Hungary. May
Lion (Panthera leo) family climbing and resting in tree. Serengeti National Park, Serengeti, Tanzania
Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean
Great egret (Ardea alba) and Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Mountain landscape reflected in lake, Basongcuo National Park, Qinghai-Tibet Plateau, Tibet, November 2011
Common raven (Corvus corax) two adults in flight, Hornoya, Norway, April
Eurasian griffon vulture (Gyps fulvus) in flight, Cevennes, France, March 2016
White fronted capuchin monkey (Cebus capucinus) feeding, March, Costa Rica
Madagascar / Malagasy kingfisher (Alcedo vintsioides) perching, Anjajavy Private Reserve, north west Madagascar
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Indian wild ass (Equus hemionus khur), group standing together, Little Rann of Kutch, Gujarat, India
Striped Dolphin (Stenella coeruleoalba) in the strait of Gibraltar, Spain
Hawfinch (Coccothraustes coccothraustes), male perched on twig, Uto, Finland, May
Bealach na Ba Pass, Wester Ross, Scotland, UK, November 2015
Namib sand snake (Psammophis namibensis) in sand dunes, Swakopmund, Erongo Region, Namibia
Honey bee (Apis mellifera) on comb with honey, Kiel, Germany, June
Rock ptarmigan (Lagopus muta) in flight, Utsjoki, Finland, April
Siskin (Carduelis spinus) male in flight preparing to land, Scotland, UK, May
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Hoopoe (Upupa epops) male in flight, Hungary May
Dwarf lionfish (Dendrochirus brachypterus) opens its awingsa (in fact enlarged pectoralDwarf lionfish (Dendrochirus brachypterus) opens its wings (in fact enlarged pectoral fins) Seraya, Tulamben area, Bali, Indonesia, Java Sea
Agave (Agave felgeri) leaves, close up. Endemic plant to Mexico. Meise National Botanic Garden of Belgium, Belgium, August
Black stork (Ciconia nigra) Hungary May
Green lacewing (Abachrysa eureka) in flight, , Tuscaloosa County, Alabama, USA Controlled conditions. May
Harlequin ladybird (Harmonia axyridis) flying, Tuscaloosa County, Alabama, USA Controlled conditions. January
Satiny Parrot snake (Leptophis depressirostris) Santa Rita, Costa Rica
Hoopoe (Upupa epops) male feeding female in courtship display, Pusztaszer, Hungary, April
Spiny predatory katydid (Listroscelis sp) photographed on a white background in mobile field studio, Osa Peninsula, Costa Rica
Clearwing moth (Cosmosoma teuthras) photographed on a white background in mobile field studio, Cordillera de Talamanca mountain range, Caribbean Slopes, Costa Rica
Shag (Phalacrocorax aristotelis) two heads showing above rock, Vardo, Norway March
Sperm whale (Physeter macrocephalus) with mouth open, Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Banded demoiselle (Calopteryx splendens) males resting on grass, Netherlands, August
Iolas blue butterfly (Iolana iolas) pair mating, Aosta Valley, Gran Paradiso National Park, Italy
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Humpback whale (Megaptera novaeangliae), mother and calf swimming just below surface in front of the West Maui Mountains. South of Lahaina, West Maui, Hawaii
Bengal tiger (Panthera tigris tigris) in forest clearing. Camera trap image. Pench National Park, Madhya Pradesh, India. January 2018
Green turtle (Chelonia mydas] swimming in open ocean, Andaman Sea, Thailand. April
Golden jackal (Canis aureus) snarling. Danube Delta, Romania. May
Hooded crown (Corvus cornix) flying with egg in beak, Danube Delta, Romania. May
Robin (Erithacus rubecula) Cadiz, Spain, March
Red deer (Cervus elaphus) stag and two hinds, Los Alcornocales Natural Park, southern Spain, September