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Southern humpback whale (Megaptera novaeangliae australis), fluke of male as he dives. Vava'u, Tonga, South Pacific
Atlantic puffin (Fratercula arctica) swimming on the ocean surface, Svalbard, Norway, June
Polar bear (Ursus maritimus) female with a single young cub, only a few months old, northern Svalbard, Norway, June
Cuttlefish (Sepia officinalis) feeding on Scorpionfish (Scorpaenidae). Tenerife, Canary Islands
Blue shark (Prionace glauca). Canary Islands. North Atlantic Ocean
Arctic fox (Alopex lagopus) following scent, in winter pelage. Svalbard, Norway. April
Arctic fox (Alopex lagopus) standing in tundra. Svalbard, Norway, July
Mute swan (Cygnus olor), two cygnets sheltering under parent's wing. Richmond Park, London, England, UK. April
Mute swan (Cygnus olor) swimming with cygnets in misty lake in morning light. Richmond Park, London, England, UK. May. Cropped image
Mute swan (Cygnus olor) cygnet swimming, cocking head. Barnes, London, England, UK. April
California condor (Gymnogyps californianus) in flight, wings radio tagged. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. 2011
Indian leopard (Panthera pardus fusca) on rock with forested hills beyond. Nilgiri Biosphere Reserve, India. Camera trap image. 2019
Honey bee (Apis mellifera) extending proboscis to nectar on Common rue (Ruta graveolens). Green aphid (Aphidoidea) beneath flower. Surrey, England, UK. July
Snowdrop (Galanthus nivalis) anthers and inner petals. Outer petals have been removedSnowdrop (Galanthus nivalis) anthers and inner petals.Outer petals have been removed. Focus stacked
Snowdrop (Galanthus nivalis) dispersing pollen by sonication which replicates buzz pollination. Controlled conditions
Chilean bellflower (Crinodendron hookerianum) releasing pollen via sonication which replicates buzz pollination. Native to Chile. Controlled conditions
New Zealand tree fuchsia (Fuchsia excorticata) with blue pollen and yellow tipped stigma. Focus stacked
Bloody bellflower (Nesocodon mauritianus), female phase with red nectar surrounding stigma. Native to Mauritius. Cultivated at Kew Gardens, London, England, UK
Morning glory (Ipomoea tricolor) flower with pollen grains scattered by visiting insects
Celandine crocus (Crocus korolkowii) carpet in area of snowmelt. Pollinated by solitary bees and bumblebees. In valley, near Ansob Pass, Tajikistan. April 2014
Common rue (Ruta graveolens) flower in visible light, nectar at base of petals. Stamens flip up sequentially with one still folded over the receptacle. Surrey, England, UK
Common rue (Ruta graveolens) flower in UV light, pollen and nectar at base of petals fluorescing. Insect pollinators are able to see position of rewards on offer. Surrey, England, UK
Tulip tree (Liriodendron tulipifera) flower, pollen and nectar fluorescing in UV light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 2/2
Porcelain orchid (Chloraea magellanica) with osmophores on lip fluorescing in UV light. Chile. Controlled conditions, focus stacked. Sequence 2/2
Chinese meadow-rue (Thalictrum delavayi) flowers. Male phase below with stigmas hanging below immature stigmas, female phase above after stamens have fallen
Amur / Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snowAmur/ Siberian tigers (Panthera tigris altaica) pair nuzzling each other in snow. Captive in tiger park, Heilongjiang Province, China. February
Cup fungus (Cookeina sp) growing on decaying wood on the rainforest floor, Corcovado National Park, Osa Peninsula, Costa Rica
Snowy owl (Bubo scandiacus) Female taking off the winter tree, Quebec, Canada
Man using pole to propel bamboo raft on Lijang / Li River, Karst peaks beyond. At dawn, Lijiang River Scenic Zone, Yangshuo county, Guangxi, China
Golden-collared Tanager (Iridosornis jelskii), in the Peruvian cloud forest. Wayqecha Biological Research Station, Peru. April. Cropped
Lapwing (Vanellus vanellus) Titchwell RSPB Reserve, Norfolk, England, UK. November
Japanese squirrel (Sciurus lis) portrait with autumn leaves, Mount Yatsugatake, Nagano Prefecture, Japan, November. Endemic species
Nazca booby (Sula granti) standing on rock at coast. Punta Suarez, Espanola Island, Galapagos. October 2015
Blue-footed booby (Sula nebouxii) colony at sunet. Cape Douglas, Fernandina Island, Galapagos
Lemon-yellow tree frog (Hyla savignyi) Cyprus. April
Mountain view towards Loch Nevis overlooking Knoydart peninsula with Skye beyond. Taken from Sgurr Cos na Breachd-laoidh, Lochaber, Scotland, UK, September 2016
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
RF - Common blue butterflies (Polyommatus icarus) resting on grass. Vealand Farm, Devon, UK. September
Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. July
RF - Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August
Red squirrel (Sciurus vulgaris) standing on log surrounded by heather, Cairngorms National Park, Scotland, UK, August
Mining bee / Little flower bee (Anthophora bimaculata) taking nectar and pollen from Common fleabane (Pulicaria dysenterica) flower, with pollen covered scopa, Surrey, England, UK. August
California sealion (Zalophus californianus) swimming. Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico
Chocolate working cocker spaniel amongst sand dunes. Formby, Merseyside, UK
Loggerhead shrike (Lanius ludovicianus) on an ice-covered fence after an ice storm, Anahuac National Wildlife Refuge, Texas, USA. January
Common loons (Gavia immer), two adults and chick swimming on a misty lake in early morning, Michigan, USA. June
Golden-fronted woodpecker (Melanerpes aurifrons), male, Cozad Ranch, Rio Grande Valley, Texas, USA, March
Roseate spoonbill (Ajaia ajaja), adult in breeding plumage flying in to land on a perch, Orlando, Florida, USA, March
