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Banded demoiselle (Calopteryx splendens) two males on grass, Netherlands
Silver washed fritillary butterfly (Argynnis paphia) female in flight with thistles, Uusimaa, Finland, August
Golden eagle (Aquila chrysaetos) portrait, captive, occurs in the Northern hemisphere
Wolf fish (Anarhichas lupus) portrait, Akuyeri, Iceland. North Atlantic Ocean
Coyote (Canis latrans) jumping in snow with its prey, Yellowstone National Park, Wyoming, USA, February
Head portrait of grey Andalusian stallion in dark barn, Northern France, Europe
Head portrait of grey Andalusian stallion running with mane flowing in arena in Northern France, Europe
Atlantic Puffin (Fratercula arctica) swimming, split level view, Skomer Island, Wales, UK, July
North Pacific right whale (Eubalaena japonica) swimming near surface, aerial view, Channel Islands National Park, California, USA. February
Orchid (Bulbophyllum sp) on white background, Botanic Garden Leiden, Netherlands, July
Marine iguana (Amblyrhynchus cristatus) two on beach, Santa Cruz Island, Galapagos
Montane Glass Frog (Centrolenella ilex) inside curled leaf. Mid-altitude rainforest, Bosque de Paz, Pacific slope, Costa Rica, Central America
Leaf-cutting bee (Megachile species) carrying leaf section. Surrey, England, UK, July
African elephants (Loxodonta africana) grazing on shore of Lake Kariba, Matusadona National Park, Zimbabwe
Green sea turtle (Chelonia mydas) Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
After an incubation period of 45 to 55 days a first hatchling Green turtle (Chelonia mydas) emerges out of the sand, Bonaire, Leeward Antilles, Caribbean region, Netherlands Antilles
Grey seal (Halichoerus grypus) large female exhales a stream of bubbles as she sleeps on shallow seaweeds (Fucus serratus) Lundy Island, Devon, UK, Bristol Channel, August
Wild Iberian lynx (Lynx pardinus) photographed with camera trap. Andalusia, Spain
Siberian crane (Grus leucogeranus) critically endangered, captive
Chamois (Rupicapra rupicapra) in deep snow trying to struggle their way out, Gran Paradiso National Park, Italy, November
Alpine landscape with snow covered trees. Gran Paradiso National Park, Italy, January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Tree Sparrow (Passer Montanus), Bayern, Germany. December
White-throated kingfisher (Halcyon smyrnensis) with Black drongo (Dicrurus macrocercus) Ranthambore National Park, Rajasthan, India
Great grey owl (Strix nebulosa) head portrait, Kuusamo, Finland, March
Japanese / Red-crowned crane (Grus japonicus) three coming into land, Hokkaido Japan February
Gannet (Morus bassanus) nearest out of focus, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Black headed gull (Chroicocephalus ridibundus) taking off, detail of legs and water droplets, Pusztaszer, Hungary, April
Beautiful pitviper (Trimeresurus venustus) juvenile, captive occurs in Thailand. Venomous species
Thornbug (Umbonia crassicornis) San Jose, Costa Rica
Cocoi heron (Ardea cocoi) resting portrait, Pantanal, Brazil
Yacare caiman (Caiman yacare) abstract view of tail and reflection at sunrise, Pantanal, Brazil
Giraffe (Giraffa camelopardalis) in evening light with rainbow, Marataba, Marakele National Park, Limpopo Province, South Africa, December
Reticulated giraffes (Giraffa camelopardalis reticulata) black and white picture of group of three, Kenya
Tail of Sperm whale (Physeter macrocephalus) diving. Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Sandhill crane (Grus canadensis) landing at sunset. North America. Highly honoured in the Art in Nature category of the Natures Best Photography Windland Smith Rice International Awards Competition
Atlantis frittilary (Speyeria atlantis) New Brunswick, Canada, July
Mazarine blue butterfly (Cyaniris semiargus) Aosta Valley, Gran Paradiso National Park, Italy
RF - Eschers blue butterfly (Polyommatus escheri) on grass, with soft focus blue flowers, Aosta Valley, Gran Paradiso National Park, Italy
RF - Monarch butterfly (Danaus plexippus) with life-cycle tied to the presence of milkweed plants (Asclepias sp). East Haddam, Connecticut, USA. September
Common blue butterfly (Polyommatus icarus) resting on Scabious (Scabiosa colombaria) flower, Volehouse Farm, Devon, UK. August 2014
RF- White water lily (Nymphaea alba) on calm water, Danube delta rewilding area, Romania
Bell glass frog (Cochranella nola) from below, photographed on a pane of glass in Manu Biosphere Reserve, Peru
Blue tit (Cyanistes caeruleus) on lichen-covered twig, England, UK May 2015
Beautiful demoiselle damselfly (Calopteryx virgo) female flying, Peerdsbos, Brasschaat, Belgium. June
Golden jackal (Canis aureus) walking on sand. Danube Delta, Romania, May
Bearded tit (Panurus biarmicus), five perched on Reed (Phragmites australis). Danube Delta, Romania. May
Golden jackal (Canis aureus) howling in grassland. Danube Delta, Romania, May
Corn bunting (Miliaria calandra), Cadiz, Andalusia, Spain, May
