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Glowworm beetle (Phengodes sp. ) male flying, Tuscaloosa County, Alabama, USA Controlled conditionsGlowworm beetle (Phengodes sp.) male flying, Tuscaloosa County, Alabama, USA Controlled conditions. July
Caribbean reef squid (Sepioteuthis sepioidea) swimming in open water at night, Guadeloupe Island, Mexico. Caribbean
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Frozen Llyn Teyrn with Y Llewydd and Mount Snowdon in background. View west from Miners Track. Snowdonia National Park, Gwynedd, Wales, UK. March 2018
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Arctic fox (Vulpes lagopus), juvenile licking lips, winter pelage. Dovrefjell National Park, Norway. September
Red bat (Lasiurus borealis) female flying; shots taken with high speed flash San Saba County, Texas, USA. Controlled conditions. July
Grey seal (Haichaoerus grypus) swimming below surface with sunbeams. Lundy Island, Devon, UK. Bristol Channel, August
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Short-finned pilot whale (Globicephala macrorhynchus) showing underside whilst vertical underwater, South Tenerife, Canary Islands, Atlantic, October
Deccan ground gecko (Cyrtodactylus albofasciatus) on forest floor, backlit with water droplets. Amboli, Maharashtra, India. Endemic to Western Ghats
Moss mimic stick insect (Trychopeplus laciniatus) showing amazing camouflage on mossy vine, photographed in tropical rainforest in mobile field studio against a white background
Strawberry poison frog (Oophaga pumilio) photographed on a white background in mobile field studio, Central Caribbean foothills, Costa Rica
Green june beetle (Cotinis nitida) in flight Williamson County, Texas, USA Controlled conditions. October
Short-finned pilot whales (Globicephala macrorhynchus) two just below surface, Los Gigantes, South Tenerife, Canary Islands, Atlantic Ocean, July
Killer whale (Orcinus orca) diving and hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Snowy egret (Egretta thula) portrait, viewed from below. St. Petersburg, Florida, USA
RF - Maidens veil / Bridal veil fungus (Phallus indusiatus) with indusium fully formed, on rainforest floor. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Warthog (Phacochoerus aethiopicus) female with piglets, Okavango Delta, Botswana
Northern raven (Corvus corax) pair perching on branch. Danube Delta, Romania, May
Golden jackal (Canis aureus) portrait. Danube Delta, Romania, May
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
Black spider monkey (Ateles chamek) portrait, captive, Peru
Tour de Goa and mountains of the Pyrenees. Casteil, Pyrenees Orientales, south west France. May 2018
Shag (Phalacrocorax aristotelis) in flight over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Marsh fritillary (Euphydryas aurinia) on fern. South of Casteil, Pyrenees Orientales, south west France. May
Scarletina bolete (Boletus luridiformis) toadstool, New Forest National Park, Hampshire, England, UK. September
Ghost glass frog (Sachatamia ilex), Costa Rica
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
Straw-coloured fruit bat (Eidolon helvum), female flying carrying pup on front. Lamin, Gambia
African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana
RF- Jaguar head profle portrait (Panthera onca) head portrait captive. Pantanal, Brazil
Polar bear (Ursus maritimus) swimming at surface of Beaufort Sea, Alaska, USA
Red squirrel (Sciurus vulgaris) stood on log in snow, Cairngorms National Park, Scotland, UK. December
Jungle cat (Felis chaus), female in search of prey. Velavadar National Park, Gujarat, India
White-naped crane (Grus vipio) flying, Inner Mongolia, China
Tea mangrove (Pelliciera rhizophorae) seed, Pochote Estuary, Costa Rica, Vulnerable species
Magnificent hummingbird (Eugenes fulgens) female, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Barn swallow (Hirundo rustica) feeding young, Monmouthshire, Wales, UK, July
Mountain Caracara (Phalcoboenus megalopterus) taking off, Laguna Canapa, Altiplano, Bolivia
RF - Mute swan (Cygnus olor) taking off from a pond Valkenhorst Nature Reserve, Valkenswaard, the Netherlands June
Mute swan (Cygnus olor) on a misty morning, with wings raised up in aggressive stance, Valkenhorst Nature Reserve, Valkenswaard, the Netherlands. December
Indian Flying Fox (Pteropus giganteus), flying, India
Stoat (Mustela erminea) looking out of hole in snow, in white winter coat, British Columbia, Canada, February
Greater roadrunner (Geococcyx californianus), adult drinking, Rio Grande Valley, South Texas, Texas, USA. June
