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Flowering rush (Butomus umbellatus) with stamens and stigma, nectar globules at base of carpels. Surrey, England, UK. Focus stacked
Christmas rose (Helleborus niger) flower, pollen on dehisced anthers and on petals fluorescing in UV light. Nectar ring appearing red. Focus stacked
Large-flowered evening primrose (Oenothera glazioviana). Bulls eye in centre of flower fluorescing in UV light guides pollinators to nectaries. At night in garden. Surrey, England, UK
Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California
Winter aconite (Eranthis hymenalis) in UV light, pollen on outer stamens that have dehisced fluorescing. Focus stacked
Tulip tree (Liriodendron tulipifera) flower with orange band to attract bees, produces copious nectar. In visible light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 1/2
White-collar burnet moth (Zygaena carniolica) nectaring on Catnip (Nepeta nuda). Greece. July
Bee beetle (Trichius fasciatus), a bee mimic, nectaring on Umbel-flowered catchfly (Silene compacta). Picks up pollen on hairy body. Russian Caucasus. June
Bar headed geese (Anser indicus), four in flight, one calling. Coming in to land at breeding site, Qinghai Lake, Qinghai Province, China
Sunlight on grassland from Qinghaia'Tibet / Qingzang railway, the worlds highest railwaySunlight on grassland from Qinghai-Tibet / Qingzang railway, the worlds highest railway. August 2009
Golden spider lily (Lycoris aurea) in rain. Yunnan Province, China. In Japan it is pollinated by swallowtail butterflies
Cremanthodium (Cremanthodium brunneopilosum). In wet meadow between Barkham and Ruo Er Gai, Sichuan Province, China
Poppywort (Meconopsis pseudointegrifolia). Yunnan Province, China
Blue tiger (Tirumala limniace) butterfly nectaring on Lantana (Lantana sp). Yunnan Province, China
Red-flanked bluetail (Tarsiger cyanurus) male front view, Jyvaskyla, Central Finland, May
Red squirrel (Sciurus vulgaris) drinking from water, Black Isle, Scotland, UK. October
Red squirrel (Sciurus vulgaris) portrait, sitting alert in autumnal woodland leaf litter, The Netherlands, November
Red squirrel (Sciurus vulgaris) alert in autumnal woodland leaf litter, The Netherlands, November
Reticulated Giraffe (Giraffa camelopardalis reticulata) standing in grassland. Laikipia, Kenya. October
Southern red-billed hornbill male (Tockus rufirostris) Mahatsu Reserve, Botswana, June
Thorny or spiny seahorse (Hippocampus histrix) portrait made off Anilao, Philippines
Tassel hyacinth (Muscari comosum) flower. Cyprus. April
Fragrant bug orchid (Anacamptis coriophora), close up. Cyprus. April
Star clover (Trifolium stellatum). Cyprus. April
Blue shark (Prionace glauca) Azores, Atlantic Ocean
Citrine wagtail (Motacilla citreola) male perched on Reed. Latvia. April
Great egret (Ardea alba) fighting over fish with Grey heron (Ardea cinerea) Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Downy emerald dragonfly (Cordulia aenea) in flight, Joutsa, Finland, July
African lion (Panthera leo) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
European robin (Erithacus rubecula), Vosges, France, December
Ring tailed lemur (Lemur catta) backlit in late afternoon light, Berenty Private Reserve, southern Madagascar
Lioness (Panthera leo) drinking at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Whooper swans (Cygnus cygnus), flying at sunset, Caerlaverock Wildfowl & Wetland Trust, Dumfries & Galloway, Scotland, November 2016
Double rainbow over Kgalagadi Transfrontier Park, Northern Cape, South Africa
Silbury hill, a Neolithic artificial chalk mound, one of the worlds largest man-made prehistoric mounds, surrounded by flowering Dandelions (Taraxacum officinale) and Rape (Brassica napus)
A Caribbean reef shark (Carcharhinus perezi) just below the surface. Split level with blue sky and clouds. Long Island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean
Culpea or Andean fox (Pseudalopex culpaeus) at rest, vicinity of Laguna Verde, Reserva Eduardo Avaroa, altiplano, Bolivia
Laysan Albatross (Phoebastria immutabilis) in nest, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Southern white admiral (Limenitis reducta) North of Lorgues, Provence, southern France, May
Banded demoiselle (Calopteryx splendens) two males on grass, Netherlands
Silver washed fritillary butterfly (Argynnis paphia) female in flight with thistles, Uusimaa, Finland, August
Golden eagle (Aquila chrysaetos) portrait, captive, occurs in the Northern hemisphere
Wolf fish (Anarhichas lupus) portrait, Akuyeri, Iceland. North Atlantic Ocean
Coyote (Canis latrans) jumping in snow with its prey, Yellowstone National Park, Wyoming, USA, February
Head portrait of grey Andalusian stallion in dark barn, Northern France, Europe
Head portrait of grey Andalusian stallion running with mane flowing in arena in Northern France, Europe
Atlantic Puffin (Fratercula arctica) swimming, split level view, Skomer Island, Wales, UK, July
