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Namib sand gecko (Pachydactylus rangei) portrait, Swakopmund, Erongo, Namibia
Eyelash palm pit viper (Bothriechis schlegelii) with tongue extended, Mindo, Pichincha, Ecuador
Werners / Tumbesian bush anole (Polychrus femoralis) on plant, Macara, Loja, Ecuador
Clarion Burrowing Owl (Athene cunicularia rostrata), Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Bateleur Eagle (Terathopius ecaudatus) female, stretching wings Kgalagadi Transfrontier Park, South Africa. January
Silk moth (Automeris zugana) sequence 2 of 2, with wings open to reveal eyespots, a means of deterring predators, photographed on a white background in mobile field studio
Red-eyed leaf frog (Agalychnis callidryas) male, Central Caribbean foothills, Costa Rica
Lemur leaf frog (Agalychnis lemur) at night, Central Caribbean foothills, Costa Rica. IUCN Red List critically endangered species
Alpine Newt (Ichthyosaura / Triturus alpestris) male in mountain pool. Nordtirol, Austrian Alps. July
Spider hunting wasp (Pompilidae) Maliau Basin, Sabah, Borneo
Red Squirrel (Sciurus vulgaris) leaping onto log, Scotland, UK, September. Highly commended in the Habitat category of the BWPA (British Wildlife Photography Awards) 2016
Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015
Grass snake (Natrix natrix) flicking tongue, Hampstead Heath, London, England, UK, August
Red Squirrel (Sciurus vulgaris) adult sitting and feeding in snowfall. Cairngorms National Park, Scotland, UK, March
RF- Serotine bat (Vespertilio serotinus) in flight above Hogweed. Captive, UK
Chaffinch (Fringilla coelebs) adult male drinking at garden pond, Scotland, UK. July
Robin (Erithacus rubecula) portrait on (Pinus sp), Black Isle, Scotland, UK March
Great Tit (Parus major) drinking at garden pond, Scotland, UK. July
Long-eared owl (Asio otus) peering round tree trunk, captive, UK
Magpie (Pica pica) perched in lichen covered tree, Titchwell, Norfolk, England, UK, February
Crested tit (Lophophanes cristatus) perched on snowy conifer branch, Scotland, March
Yellow Hammer (Emberiza citrinella) UK, March
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Chocolate-dip chromis (Chromis dimidiata) male over out of focus hard coral (Porites nodifera). Sha ab Claudia, Fury Shoal, Egypt. Red Sea
RF - Split level photo of a Whale shark (Rhincodon typus) feeding at the surface. Isla Mujeres, Yucatan Mexico. Caribbean Sea
A stand of Pillar coral (Dendrogyra cylindrus) growing on a coral reef, East End, Grand Cayman, Cayman Islands
RF- Peregrine Falcon (Falco peregrinus) landing on moorland with its wings spread. Scotland, UK, March
Irish march orchid (Dactylorhiza majalis) in flower, Sainte Marguerite, France, May
RF- Sunset on Kimmeridge Bay, Isle of Purbeck, Jurassic Coast World Heritage Site, Dorset, England, UK. November 2008
RF- Little blue heron (Egretta caerulea) and Great egret (Ardea alba) in lagoon at sunrise. Fort Myers beach, Gulf Coast, Florida, USA, March
RF- Two Greater flamingos (Phoenicopterus ruber) flying across sunset sky, Namibia
Common / Eurasian cranes (Grus grus) taking flight for roasting site, at sunset, silhouetted. Lake Hornborga, Sweden, April
RF - Marbled White butterfly (Melanargia galathea) resting on common spotted orchid, Dunsdon Nature Reserve, Devon, UK. July
Two Great egrets (Ardea alba) wading, silhouetted at dawn, Keoladeo National Park, Bharatpur, Rajasthan, India
Great Tit (Parus major) perched on branch with yellow from autumn trees in the background. Southern Norway, October
Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Snow petrel (Pagodroma nivea) flying, Antarctica, November
Sandy beach and bright blue ocean, Fuerteventura, Canary Islands. April 2013
Male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, Ras Katy, Sharm El Sheikh, Sinai, Egypt. Gulf of Aqaba, Red Sea
RF- Small jellyfish (Pelagia noctiluca) swimming beneath surface. Marine Protected Area of Portofino (Area Marina Protetta, Portofino), Liguria, Italy. Mediterranean Sea
Masai giraffes (Giraffa camelopardalis tippelskirchi) in dry season, Masai-Mara Game Reserve, Kenya. March
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Artic fox (Vulpes lagopus) portrait, captive, Norway, February
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
Ancient dead Camelthorn tree (Vachellia erioloba) trees with red dunes and mist, Namib desert, Deadvlei, Sossusvlei, Namibia. August 2015
Gentoo penguins (Pygoscelis papua) calling, Cuverville Island. Antarctic Peninsula, Antarctica
Gentoo Penguin (Pygoscelis papua) Wiencke Island, Antarctic Peninsula, Antarctica
Masai giraffe (Giraffa camelopardalis tippelskirchi) two silhouetted at sunrise, Msai Mara Game Reserve, Kenya
