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Golden jackal (Canis aureus) howling in grassland. Danube Delta, Romania, May
Corn bunting (Miliaria calandra), Cadiz, Andalusia, Spain, May
Corn bunting (Emberiza calandra) singing, Prado del rey, Cadiz, Spain, March
Western clubtail dragonfly (Gomphus pulchellus), Sierra de Grazalema Natural Park, southern Spain. June
Great grey shrike (Lanius meridionalis) perched on a branch of Montpellier maple (Acer monspessulanum). Sierra de Grazalema Natural Park. southern Spain. December
Orchid (Orchis olbiensis), Sierra de Grazalema Natural Park, Benaocaz, southern Spain, May
Arctic fox ( Vulpes lagopus) in snow carrying snowgoose egg, Wrangel Island, Far East Russia, May
Emperor penguin (Aptenodytes forsteri) chick, Ross Sea, Antarctica
Emperor penguin (Aptenodytes forsteri) walking, Gould Bay, Weddell Sea, Antarctica
Head portrait of grey Andalusian stallion in dark barn. Northern France, Europe
Indian rhinoceros (Rhinoceros unicornis) female. Kaziranga National Park, India
Ebony jewelwing (Calopteryx maculata) New Brunswick, Canada, July
Long-tailed manakin (Chiroxiphia linearis) perched on lek (display perch), Costa Rica
European red fox (Vulpes vulpes crucigera) in deep snow in front of steep rocks. Gran Paradiso National Park, Italy. December Highly commended in the Portfolio category of the Terre Sauvage Nature
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Banded demoiselle (Calopteryx splendens) male resting on water lily, River Wensum, Norfolk, England, UK, July
Garden bumblebee (Bombus hortorum), flying to Foxglove (Digitalis purpurea), flower, Monmouthshire, Wales, UK. June
RF - Mallard (Anas platyrhynchos) female swimming with with young chicks following, Valkenhorst Nature Reserve, The Netherlands June
Cheetah (Acinonyx jubatus) male on the hunt with Crater rim in the background, Ngorongoro Crater, Tanzania
Plateau pika (Ochotona curzoniae), Tibetan Plateau, Qinghai, China
Wild yak herd (Bos mutus) Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
Common blue butterfly (Polyommatus icarus) resting on devils bit scabious (Succisa pratensis), Vealand Farm, Devon, UK. August
Snowy owl (Bubo scandiacus) Female taking off the winter tree, Quebec, Canada
Roe deer (Capreolus capreolus) entering pond about to swim across it. Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. November
Robin (Erithacus rubecula) taking off from snow, Hampstead Heath, London, UK, January
Four Mute swans (Cygnus olor) walking over snow, Hazerswoude, The Netherlands, February
European brown bear (Ursus arctos) resting in the snow, Finland, April
Red squirrel (Sciurus vulgaris) standing on hind legs, Brasschaat, Belgium, February
Meadow brown butterfly (Maniola jurtina) on Common poppy seedhead (Papaver rhoeas), De Inslag, Brasschaat, Belgium August
Red-tailed Tropicbird (Phaethon rubricauda) flying portrait, Nosy Ve, Madagascar
RF- (Pinus sylvestris) at sunrise, Klein Schietveld, Brasschaat, Belgium
RF -Three Bison (Bison bison) walking through snow with herd feeding in background. Yellowstone National Park, Wyoming, USA. January
Coyote (Canis latrans) in snow, Yellowstone National Park, Wyoming, USA, February
Grizzly cubs (Ursus arctos) two play fighting in the mist, near Brooks Falls in Katmai National Park, Alaska, USA, July
RF - Coyote (Canis latrans) in snow, Yellowstone. February
Coyote (Canis latrans) concealed in the snow, Yellowstone National Park, Wyoming, USA. January
Red fox (Vulpes vulpes) in snow, Yellowstone National Park, Wyoming, USA. Wyoming, USA, January
Western Scrub-Jay (Aphelocoma californica), adult calling on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Horned puffins (Fratercula corniculata) pair interacting by touching bills while perched on cliff ledge, St. Paul Island, Pribilofs, Alaska, USA, July
Burrowing owl (Athene cunicularia) Adams County, Washington, USA. May
RF- Red squirrel (Sciurus vulgaris) portrait, in snow, Cairngorms National Park, Scotland, March 2007
RF- Solar halo over the Laitaure delta, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden. September 2008
RF- White-tailed Sea Eagle (Haliaeetus albicilla) turning in flight, Flatanger, Norway, August
RF- Red squirrel (Sciurus vulgaris) jumping. Cairngorms National Park, Scotland, March 2012
RF- Two Small red damselflies (Ceriagrion tenellum) covered in morning dew. Arne (RSPB) Nature Reserve, Dorset, UK. August
RF- Rock ptarmigan (Lagopus mutus) standing on boulder, camouflaged against snow in winter plumage. Cairngorms National Park, Highlands, Scotland, UK, February
Red squirrel (Sciurus vulgaris) with autumn colours, Cairngorms National Park, Highlands, Scotland, UK. October
Red squirrel (Sciurus vulgaris) climbing down Scots pine tree, with heather in bloom, Highlands, Scotland, UK, August
