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Scarlet Macaw (Ara macao) couple in flight and breaking to land Corcovado National Park, Osa peninsula, Costa Rica
Large whip goby (Bryaninops amplus) with a parasitic copepod (with two trailing egg ribbons) perched on Red whip coral. Whale Rock. Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia
Yellow mongoose (Cynictis penicillata), two pups nose to nose. Kalahari Gemsbok National Park, Kgalagadi Transfrontier National Park, Northern Cape, South Africa
RF - Chinstrap penguins (Pygoscelis antarctica) displaying, Half-Moon Island, South Shetland Islands, Antarctica (This image may be licensed either as rights managed or royalty free.)
Roe deer (Capreolus capreolus) on a misty morning at the edge of a pond. Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. December
Mute swan (Cygnus olor) on river in evening light, Gelderse Poort, near Nijmegen, The Netherlands. June
Egyptian goose (Alopochen aegyptiaca) in flight, London, UK, February
Goldeneye (Bucephala clangula) male and two females on woodland loch at dawn on misty morning, Loch Vaa, Cairngorms National Park, Scotland, UK, April
RF- Whooper Swan (Cygnus cygnus) standing on frozen loch at dawn in winter. Loch Insh, Cairngorms National Park, Scotland, UK
Caribbean flamingo (Phoenicopterus ruber) landing, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Lemon-yellow tree frog (Hyla savignyi) peering out from behind plant. April
RF - Adelie penguins (Pygoscelis adeliae) on blue ice berg. Graham Passage, Antarctic Peninsula, Antarctica. (This image may be licensed either as rights managed or royalty free.)
Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
Grey wolf (Canis lupus) standing among trees in snow-covered forest, Finland. November
Caribbean flamingo (Phoenicopterus ruber) feeding in waves, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August. Bookplate
Two Great egrets (Ardea alba) face to face against white sky, Lake Csaj, Pusztaszer, Hungary. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015
Caribbean flamingos (Phoenicopterus ruber) line of adults standing in water, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
Broadhaven Beach, Pembrokeshire Coast National Park, Pembrokeshire, Wales, March 2012
Caribbean flamingo (Phoenicopterus ruber) flock in distance at dusk, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, August
Caribbean flamingos (Phoenicopterus ruber) silhouetted at dawn, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August
Puffin (Fratercula arctica) through soft focus vegetation. Great Saltee, Saltee Islands, County Wexford, Ireland, June
Bee orchid (Ophrys apifera), Powerstock Common, Dorset Wildlife Trust, Dorset, England, UK. June
Green sea turtle (Chelonia mydas) swimming over sand ripples in shallow water, Eleuthera, Bahamas. Endangered
Arctic fox (Alopex lagopus) in winter coat, walking across snow, Svalbard, Norway. April
Ringed seal (Pusa hispida) resting on ice, Svalbard, Norway. April
Ptarmigan (Lagopus mutus) in winter plumage, resting in snow, Svalbard, Norway. April
Yellow-bellied toad (Bombina variegata) male in pond, calling, Lucerne, Switzerland. July
Red eyed tree frog (Agalychnis callidryas) resting on tree trunk at night, Calakmul Biosphere Reserve, Campeche, Mexico. Cropped
Wine throated hummingbird (Selasphorus ellioti) male, courtship display, Cloud forest, Atitlan Volcano, Solola, Guatemala
Little bittern (Ixobrychus minutus) perched in reedbed calling, Valkenhorst nature reserve, Valkenswaard, the Netherlands. May
Nilgiri tahr (Nilgiritragus hylocrius) standing next to road signage meant to warn vehicles about the possible road crossings of tahrs in the area, Valparai, Tamil Nadu, India
Crimson-rumped toucanet (Aulacorhynchus haematopygus) peering our from nest hole in tree trunk, Pichincha, Ecuador
Rufescent tiger heron (Tigrisoma lineatum) portrait, Yasuni National Park, Orellana, Ecuador
Black-mantled tamarin (Leontocebus nigricollis) sitting on branch in rainforest, Cuyabeno Reserve, Sucumbios, Ecuador
Andean cock-of-the-rock (Rupicola peruvianus) perched on branch in rainforest, Sumaco National Park, Napo, Ecuador
Puma (Puma concolor) female with two cubs, Torres del Paine National Park, Magallanes, Chile
Two Pallas's cat (Otocolobus manul) kittens waiting at their den for their mother to return from hunting, Mongolia, June, 2017. Filmed for BBC series Big Cats
A female Pallas's cat (Otocolobus manul) with two young kittens hiding amongst rocks at their den site, Mongolia, June, 2017. Filmed for BBC series Big Cats
Lionfish (Pterois volitans) patrolling the edge of Suzie's Bommie, a healthy reef and popular dive site accessible from Lololata Island Resort near Port Moresby, Coral Sea, Papua New Guinea
Humpback whale (Megaptera novaeangliae) rear view of males escort, with adult female and male calf in the background. Vava'u, Tonga, South Pacific
Polar bear (Ursus maritimus) very young cub, probably only a few months old, standing up to take a look around, northern Svalbard, Norway, June
Atlantic puffin (Fratercula arctica) head profile, Iceland, June
Kestrel (Falco tinnunculus) male, perched on fallen branch, UK. May
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Hooded crow (Corvus corone cornix) standing on water-covered ice, with reflection of buildings at sunset in background, Helsinki, Finland. December
Sperm whale (Physeter macrocephalus) tail slapping, Dominica, Caribbean Sea, Atlantic Ocean. Photo taken under permit. January
Egyptian goose (Alopochen aegyptiaca) gosling swimming in the water, London, UK, February