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Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
American bison (Bison bison) Yellowstone National Park, Montana, USA
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Black-throated Diver (Gavia arctica) Rokua, Finland, May
European hare (Lepus europaeus), Wirral, England, UK, May
Jaguar (Panthera onca) male walking, Pantanal, Brazil
Scarlet macaws (Ara macao) La Selva, Costa Rica
Silhouette of Common peafowl (Pavo cristatus) perched in a tree at dawn. Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Arctic Fox (Alopex / Vulpes lagopus) yawning, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Japanese cranes (Grus japonensis) displaying in snow, Hokkiado, Japan, February
Willow Grouse (Lagopus lagopus) male perched on snow, Finland, March. Bookplate from Danny Greens The Long Journey North
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Stellers sea-eagle (Haliaeetus pelagicus) landing on pack ice, Hokkaido, Japan, February
Elephant hawk moth (Deilephila elpenor) portrait, Brackagh Moss National Nature Reserve, Portadown, County Armagh, Northern Ireland. April
Small pearl-bordered fritillary butterfly (Boloria selene), Marsland mouth, Cornwall, UK. JUne 2016
Pine marten (Martes martes) reflected in water, Ardnamurchan Peninsula, west coast of Scotland, UK. Highly commended in the Mammals category of the British Wildlife Photography Awards (BWPA)
Weasel (Mustela nivalis) looking out of hole on woodland floor with snowdrops, Sheffield, England, UK
Young urban Red fox (Vulpes vulpes). Bristol, UK. August
RF- Goldfinch (Carduelis carduelis) reflected in pool, Worcestershire. November
Barn owl (Tyto alba) in flight, captive, Cumbria, UK
A portrait of a Mediterranean parrotfish (Sparisoma cretense) above an algae covered rock. El Hierro, Canary Islands, Spain. East Atlantic Ocean
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February
Edwards Atlas Moth (Attacus edwardsii) in defensive posture, Bhutan, June
Siberian musk deer (Moschus moschiferus) male in snow, Irkutsk, Russia. January
Bornean horned frog (Megophrys nasuta) on forest floor. Danum Valley, Sabah, Borneo, Malaysia
Ariel toucan (Ramphastos ariel) perched on branch, Atlantic Rainforest, Rio de Janeiro city, Rio de Janeiro State, Brazil
Emperor penguin (Aptenodytes forsteri), chick in brood pouch of parent, Snow Hill Island, Antarctic Peninsula
Pied avocet (Recurvirostra avosetta) feeding in shallow water at dawn, Etosha National Park, Namibia
Great crested grebe (Podiceps cristatus) carrying chick on back, The Netherlands, May
Giant panda cub (Ailuropoda melanoleuca) portrait Yuan Meng, first giant panda ever born in France, age 10 months. Captive at Beauval Zoo, Saint Aignan sur Cher, France
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Imbabura tree frog (Hypsiboas picturatus) portrait, Canande, Esmeraldas, Ecuador
Tropical shield mantis (Choeradodis sp. ) (Possibly C. rhombicollis) Canande, EsmeraldasTropical shield mantis (Choeradodis sp.) (Possibly C. rhombicollis) Canande, Esmeraldas, Ecuador
African elephant (Loxodonta africana) herd walking in the plains during the dry season, Masai-Mara Game Reserve, Kenya. October
Vogelkop superb bird-of-paradise (Lophorina niedda) male, performing courtship display on his display log, flashing false blue eye spots, a bright yellow gape, and blue breast shield
Green tree ants (Oecophylla smaragdina) defending their leaf nest, Daintree River, Wet Tropics World Heritage area, Queensland, Australia
Wren (Troglodytes troglodytes) perched amongst autumn foliage. Uto, Finland. October
Portuguese man o war (Physalia physalis) Tenerife, Canary Islands
Great tit (Parus major) perched on branch, ready to fly, Moselle, France, February
Green turtle (Chelonia mydas) feeding on on Turtlegrass (Thalassia testudinum) seagrass bed. The Bahamas
Black-crowned Central American squirrel monkey (Saimiri oerstedii) pair close-up Manuel Antonio National Park, Quepos, Costa Rica
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
RF - Lion (Panthera leo) male on savanna with dramatic storm clouds, thundery sky and light rays in the background. Ngorongoro Crater, Tanzania. Composite image
