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Portrait of a Black-capped chickadee, (Poecile atricapillus) with white background, Block island, Rhode Island, USA. Bird caught during scientific research
Eurasian crag martin (Ptyonoprogne rupestris) Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China. May
Grey heron (Ardea cinerea) wading at night, reflected in water. Lincolnshire, UK, September
RF - White-legged damselfly (Platycnemis pennipes) peering over Oxeye daisy (Leucanthemum vulgare) flower. The Netherlands. August
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Pair of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Red fox (Vulpes vulpes) in snowy landscape with trees and two Crows (Corfus corone) winter, Jura, Switzerland
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Resplendent quetzal (Pharomachrus mocinno) male, Talamanca mountains, Costa Rica
Small Tortoiseshell Butterfly (Aglais urticae) feeding on scabious flowers in hay meadow, Wensum Valley, Norfolk, England, UK, July
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Sumatran tiger (Panthera tigris sondaica). Captive, with digitally added leaf pattern
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Coyote (Canis latrans) walking through deep winter snow. Yellowstone National Park, Wyoming, USA. January
Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March
Lammergeier / Bearded vulture (Gypaetus barbatus) in flight with the out-of-focus moon behind. Ladakh, Himalayas, northern India
Mother of pearl (Pleuroptya ruralis) moth nectaring on Ragwort (Jacobaea vulgaris) at night. Surrey, England, UK. July
Garden bumblebee (Bombus hortorum) queen nectaring on Comfrey (Symphytum Hidcote Pink ) flowers. In garden, Surrey, England, UK. March
Lined seahorses (Hippocampus erectus) amongst corals, The Bahamas
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
Red squirrel (Sciurus vulgaris) sitting on fallen tree in pine woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August
Gannet (Morus bassanus) fluffy chick with its parent. Shetland Islands, Scotland, UK, August
American bison (Bison bison) male walking through deep snow. Hayden Valley. Yellowstone National Park, Wyoming, USA. January
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Robin (Erithacus rubecula) feeding on Gelder Rose berries, Germany. January, January
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Zanzibar red colobus monkey (Procolobus kirkii) crossing the main road through the Reserve, Jozani forest, Jozani Chwaka Bay NP, Zanzibar, Tanzania, August
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Bearded pig (Sus barbatus) digging in sand, foraging for crabs on beach, Bako National Park, Sarawak, Borneo
Grey whale (Eschrichtius robustus) tail, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Coal tit (Periparus ater) on mossy log with reflection, England, UK. January
Ladder-backed Woodpecker (Picoides scalaris), adult male perched on icy branch of Possum Haw Holly (Ilex decidua) with berries, Hill Country, Texas, USA. February
Violet-tailed sylph hummingbird (Aglaiocercus coelestis) Mindo, Pichincha, Ecuador
White prominent moth (Leucodonta bicoloria) County Kerry, Ireland
White ibis (Eudocimus albus) group of four in flight above clouds, Fort Myers Beach, Gulf Coast, Florida, USA, March
RF- Mute swan (Cygnus olor) on water in winter dawn mist. Loch Insh, Cairngorms National Park, Highlands, Scotland UK, December
Red squirrel (Sciurus vulgaris) gnawing red deer antler for minerals, Cairngorms National Park, Highlands, Scotland, UK, June
Red squirrels (Sciurus vulgaris) interacting, Cairngorms National Park, Highlands, Scotland, UK, January
Hen Harrier (Circus cyaneus) adult female in flight over heather moorland, Scotland, UK. July
King cobra (Ophiophagus hannah) in strike pose, Malaysia
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
