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Red fox (Vulpes vulpes) resting in the snow, Churchill, Cananda, November
Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February
Yellow-plumed parakeets (Leptosittaca branickii) at nest cavity in wax palm stump, Tapichalaca Biological Reserve, Ecuador
Barn owl on post in flooded Somerset Levels. England, UK, Europe
Bateleur eagle (Terathopius ecaudatus) head portrait, Captive, occurs in Africa
Red squirrel (Sciurus vulgaris) having a drink, Black Isle, Scotland, UK. October
Long-eared owl (Asio otus) in a large tree, Cheshire, UK, March
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Wren (Troglodytes troglodytes) on frost covered fern, Dunham Massey, Cheshire, UK. Winner of the Habitat Category of the British Wildlife Photographer of the Year Awards (BWPA) 2017
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Desmarests hutia (Capromys pilorides doceleguas), Jardines de la Reina / Gardens of the Queen, Caribbean Sea, Ciego de Avila, Cuba, January
Himalayan monal (Lophophorus impejanus) male perched on branch, Uttarakhand, India
Golden eagle (Aquila chrysaetus) Kuusamo, Finland, January
Coal tit (Periparus ater) on a branch with lichen, Vendee, France, December
Portrait of a Green sea turtle (Chelonia mydas) in morning light. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Two Sulawesi black macaques / Celebes crested macaques (Macaca nigra) juveniles, in woodland, portrait, Tangkoko National Park, northern Sulawesi, Indonesia. Critically endangered
Golden plover (Pluvialis apricaria) chick camouflaged in ground nest, Norway June
Three Red-crowned cranes (Grus japonensis) in flight. Hokkaido, Japan. March
Mimika bobtail squid (Euprymna morsei), Hokkaido, Japan, Pacific Ocean
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Pelagic thresher shark (Alopias pelagicus) swimming in open ocean, Malapascua Island, Philippines, Pacific Ocean
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
Bison (Bison bison) female with calf walking through snow in front of frost-covered forest, Yellowstone National Park, USA
Caspian Tern (Sterna caspia) in flight, diving for prey. Lake Ellesmere, Canterbury, South Island, New Zealand
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
White tailed eagle (Haliaeetus albicilla) calling in flight, Isle of Mull, Argyll Scotland, UK. October
Giant pacific octopus (Enteroctopus dofleini) swimming freely after release from captivity, Vancouver Island, British Columbia, Canada, Pacific Ocean
Spotted eagle rays (Aetobatus narinari) swimming, Hawaii, USA
Common buzzard (Buteo buteo) in flight with hoar frost. Northamptonshire, UK, January
Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Bottlenose dolphins (Tursiops truncatus) at the surface at sunrise. Kwazulu-Natal, South Africa
Common blue butterfly (Polyommatus icarus) resting on Devil's bit scabious (Succisa pratensis), Volehouse Farm, Devon, UK. August
Golden eagle (Aquila chrysaetus), Kuusamo, Finland, January
Lammergeier / Bearded vulture (Gypaetus barbatus) in flight with the out-of-focus moon behind. Ladakh, Himalayas, northern India
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
Brown bear (Ursus arctos arctos) female and two cubs, aged 1 year, emerging from den in winter, Bavarian Forest, Germany, captive, January
1468136 - Mandrill male (Mandrillus sphinx) abstract of brightly coloured ischial1468136 - - Mandrill male (Mandrillus sphinx) abstract of brightly coloured ischial callosity on buttocks, Lekedi National park, Gabon
Green hairstreak butterfly (Callophrys rubi) on hawthorn leaf, Wiltshire, England, UK, April
Aerial view of Reindeer (Rangifer tarandus) fording water whilst being herded by Nenet herders. Nenets Autonomous Okrug, Arctic, Russia, July 2017
RF- Carrion crow (Corvus corone) flying from a snow covered pine tree in a winter landscape. Valsavarenche, Gran Paradiso NP, Italy, March
Clarks Grebe (Aechmophorus clarkii) with yawning chick riding on back, Bear River Refuge, Utah, July
Ptarmigan (Lagopus muta) male on snowy ridge on Cairngorm mountain, Cairngorms National Park, Scotland, UK, March
White-throated dipper (Cinclus cinclus) winter with river flooding, Kuusa, Laukaa, Keski-Suomi, Lansi- ja Sisa-Suomi / Central and Western Finland, Finland. January
