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RF - Keel-billed toucan (Ramphastos sulfuratus) perched on mossy branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)
Borneo giant forest scorpion (Heterometrus longimanus) resting inside a fallen hollow log. Danum Valley, Sabah, Borneo. Photographed with natural light and UV light
Male Fossa (Cryptoprocta ferox) climbing down tree trunk from forest canopy. Mid-altitude rainforest, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Giant anteater (Myrmecophaga tridactyla) Serra de Canastra National Park, Brazil
Barn Swallow (Hirundo rustica) in flight. Portugal, Europe
Red deer (Cervus elaphus) stag in upland landscape. Lochcarron, Highlands, Scotland, UK
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Small pearl-bordered fritillary (Boloria selene) butterfly on oxeye daisy (Leucanthemum vulgare), Marsland mouth, North Devon, UK. May 2017
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Head portrait of grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
Head portrait of black Merens stallion with long mane running in pasture. Northern France, Europe. February
Osprey (Pandion haliaetus) in flight catching a fish, Finland, July
Raccoon (Procyon lotor) peering out from behind tree trunk, Baxter State Park, Maine, USA
Yellow-headed Picathartes / White necked rockfowl (Picathartes gymnocephalus) Kambui Hills, Sierra Leone, Vulnerable species. Photograph taken on location for BBC Africa series, September 2010
Wild female Amur leopard (Panthera pardus orientalis) on rocky hillside, Kedrovaya Pad reserve, Primorsky Krai, Far East Russia, January. Critically endangered species
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
Giant Panda cub (Ailuropoda melanoleuca) climbing. Yuan Meng, first Giant panda even born in FranceGiant Panda cub (Ailuropoda melanoleuca) climbing.Yuan Meng, first Giant panda even born in France, is now aged 8 months. and likes very much to stay in trees, Beauval Zoo, France
Eastern chipmunk (Tamias striatus) gathering chestnuts, Chestnut Hill, Philadelphia, Pennsylvania, USA, September
Stellers sea-eagle (Haliaeetus pelagicus) in flight, Hokkaido, Japan, February
Water fleas (Daphnia sp. ) and a green algae (Volvox aureus) in water from a garden pondWater fleas (Daphnia sp.) and a green algae (Volvox aureus) in water from a garden pond. Derbyshire, UK. September. Digital composite
Fen raft spider / Great raft spider (Dolomedes plantarius) sub-adult. Norfolk Broads, UK, September. Vulnerable species
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Angular roughshark (Oxynotus centrina) a deepsea species living at 80-300m depth, caught by fishermen and released, Costa Brava, Catalunya, Spain, Mediterranean Sea
Plains zebra (Equus quagga) grooming foal, Kruger National Park, South Africa, July
Giant panda cub (Ailuropoda melanoleuca) portrait Yuan Meng, first giant panda ever born in France, age 10 months. Captive at Beauval Zoo, Saint Aignan sur Cher, France
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Red deer (Cervus elaphus) backlit at dawn with visible breath. UK. October
Flock of ducks silhouetted against sunset flying over field in winter in the Uitkerkse Polder nature reserve near Blankenberge, West Flanders, Belgium. February 2020
Snow leopard (Panthera uncia) female in snow, Hemis National Park, Ladakh, India
Goldenrod crab spider (Misumena vatia) yellow female on Honesty flower, Bristol, UK, April
Great Spotted Woodpecker in snow (Dendrocopos major), Scotland, February
Brown bear (Ursus arctos) male, portrait. Martinselkonen, Kainuu, Finland. June
Bamboo lemur (Hapalmur griseus) in tree. Andasibe-Mantadia National Park, Madagascar
Short-beaked echidna (Tachyglossus aculeatus). Francois Peron National Park, Shark Bay, Western Australia. October
Japanese crane (Grus japonensis) displaying, wings in heart shape, Hokkiado, Japan, February
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Sable (Martes zibellina) in tree, Irkutsk, Siberia, Russia. November
Roughtail stingray (Dasyatis centura) on sea floor, Tenerife, Canary Islands
Red fox (Vulpes vulpes) in winter snow, Jura, Switzerland
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Eastern tiger salamander (Ambystoma tigrinum) North Florida, USA. December