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Emperor penguin (Aptenodytes forsteri) viewed through hole in iceberg at Snow Hill Island rookery, Antarctica. November
Eurasian flamingos (Phoenicopterus roseus) lined up on Pulicat Lake, Tamil Nadu, India, January 2013
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
Puffin (Fratercula arctica) standing on flowering Sea thrift (Armeria maritima). Portrait. Fair Island, Shetland Islands, Scotland, July
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Barn owl (Tyto alba) in flight. Suffolk, UK. June
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Sea otter (Enhydra Lutris) with forelegs raised, Sitka Sound, Alaska, USA, August
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Hermit thrush (Catharus guttatus) feeding on blue juniper berry (cone) of Eastern Red-Cedar (Juniperus virginiana) in mid-November, Freeville, New York, USA
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
European roller (Coracias garrulus) in flight, Pusztaszer, Hungary, May
American crocodile (Crocodylus acutus) shows off its teeth. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Acorn weevil (Curculio glandium) Portrait on an Oak bud, Hertfordshire, England, UK, May - Focus Stacked - Captive
Arctic fox (Vulpes lagopus), juvenile looking at camera, portrait, winter pelage. Dovrefjell National Park, Norway. September
Five white Camargue horses running through the water in Southern France, Europe. May
Common kestrel (Falco tinnunculus) perched on a branch, Valencia, Spain, February
Malaysian Orchid Mantis (Hymenopus coronatus) pink colour morph, camouflaged on an orchid. Captive, from Malaysia
Splendid leaf frog (Cruziohyla calcarifer) La Selva Field Station, Costa Rica
Grey Wolf (Canis lupus), in forest, captive, USA
Close up portrait of the face of a Nurse shark (Ginglymostoma cirratum) resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas
Cuckoo (Cuculus canorus) perched on a branch. Thursley Common, Surrey, UK, May
Polar bear cubs (Ursus maritimus) triplets age 2-3 months next to their mother. Wapusk National Park, Manitoba, Canada. February
Portrait of a Mandarin duck (Aix sponsa) male animal and female. UK. Introduced species
Peregrine falcon (Falco peregrinus) in flight over cotton grass flowers, Vaala, Finland, June
Siberian flying squirrel (Pteromys volans orii) flying directly over photographer. Hokkaido, Japan. February
Trap-jaw ant (Odontomachus chelifer) portrait, Wayqecha, Peru
Panther chameleon (Furcifer pardalis) sitting on branch with black background, Nosy Faly, Madagascar
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
Mating pair of chalkhill blue butterflies (Lysandra coridon) with wings closed resting on Devils-bit scabious (Succisa pratensis), Hatch Hill, Somerset, UK. August
Bald eagle (Haliaeetus leucocephalus) pair flying with claws linked during courtship flight, Homer, Alaska, USA, March
Marine iguana (Amblyrhynchus cristatus) swimming underwater, Fernandina Island, Galapagos
RF- Jaguar (Panthera onca) head portrait, captive
DELETE DUPLICATE - RF - Wild buckskin Mustang stallion walking towards waterhole on dusty dry summer day, Sand Wash Basin, Colorado, USA. August
Portrait of a Golden snub-nosed monkey (Rhinopithecus roxellana) screaming and showing its teeth, Foping Nature Reserve, Shaanxi, China. Endangered species
Malaysian Orchid Mantis (Hymenopus coronatus) white colour morph camouflaged on an orchid. Captive. Originating from Malaysia
Indian peafowl (Pavo cristatus) peacock displaying feathers, captive, occurs in South Asia
RF - Arctic Fox (Vulpes lagopus) portrait in winter coat, Svalbard, Norway, April
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Pod of Indian Ocean bottlenose dolphin (Tursiops adunctus) swim over a coral reef. Sha ab El Erg, Hurghada, Egypt. Red Sea
RF - Composite image of tropical nudibranchs on a black background showing variety and abundance of nudibranch species, Indo-Pacific
Deep sea jellyfish (Periphylla periphylla) juvenile, Trondheimsfjord, Norway, Atlantic ocean
RF - Keel-billed toucan (Ramphastos sulfuratus) perched on mossy branch. Boca Tapada, Costa Rica. (This image may be licensed either as rights managed or royalty free.)
Borneo giant forest scorpion (Heterometrus longimanus) resting inside a fallen hollow log. Danum Valley, Sabah, Borneo. Photographed with natural light and UV light
Male Fossa (Cryptoprocta ferox) climbing down tree trunk from forest canopy. Mid-altitude rainforest, Andasibe-Mantadia National Park, eastern Madagascar. Endangered
