Animal Behaviour Collection (page 16)

Australasian gannet / Takapu (Morus serrator) breeding colony on cliffs, Muriwai gannet colony, Auckland, North Island, New Zealand, June

Munks mobula ray / Devilray (Mobula munkiana) leaping out of the water, Sea of Cortez, Gulf of California, Baja California, Mexico, February

Two rainbow lorikeets (Trichoglossus moluccanus) on branch, Werribee, Victoria, Australia

Little bittern (Ixobrychus minutus) watching Great warbler (Acrocephalus arundinaceus) with prey, Lake Csaj, Kiskunsagi National Park, Hungary

Corncrake (Crex Crex) calling, with breath vapor in early morning, Kiskunsagi National Park, Hungary

Montezuma oropendola (Psarocolius montezuma) hanging upside down as part of a courtship display. Costa Rica

Black-backed jackals (Canis mesomelas) two fighting with Plains / Burchell zebra (Equus burchelli) in foreground, Kalahari Desert, Nxai Pans National Park. Botswana

Killer whale (Orcinus orca) hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April

Reef manta (Manta alfredi) mouth open feeding at night, South Ari atoll, Maldives, Indian Ocean

Killer whale (Orcinus orca) hunting inside herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April

Spirit / Kermode bear (Ursus americanus kermodei) catching fish in river, most likely salmon, the Great Bear Rainforest, British Columbia, Canada, October

White tailed sea eagle (Haliaeetus albicilla) interacting with Great egret (Ardea alba) Lake Csaj, Kiskunsagi National Park, Hungary. January

Western diamond-backed rattlesnake (Crotalus atrox) Texas, USA, April

Great egret (Ardea alba) and Grey heron (Ardea cinerea) squabbling, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January

Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species

Long-tailed tit (Aegithalos caudatus caudatus) placing feather in nest, Finland. April

Blackcap (Sylvia atricapilla) female and two males huddling together for warmth before going to sleep, Finland, May

Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March

Afrian lioness (Panthera leo) playing with her juvenile cub aged 2 years in Duba Plains concession. Okavango delta, Botswana

Young ground squirrels (Xerus inauris) fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January

Common dolphin (Delphinus delphis) close up of dolphin surfing at speed, Hebrides, Scotland, UK

Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa

Bluethroat (Luscinia svecica) singing on a branch Vendee, France, May

Clarion Lizard (Urosaurus clarionensis) displaying, IUCN Vulnerable, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site

Seychelles Sunbird (Cinnyris dussumieri), adult male singing, Praslin Island, Republic of Seychelles

Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela

Common midwife toad (Alytes obstetricans), male toad carrying eggs, Vaucluse, France

Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname

Striped Dolphin (Stenella coeruleoalba) in the strait of Gibraltar, Spain

Hippopotamus (Hippopotamus Amphibius) territorial aggressive behavior, Chobe National Park, Botswana

Scorpion (Heterometrus sp) fluorescing in ultraviolet light at night on forest floor, Danum Valley, Sabah, Borneo

Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea

Flame anthias (Pseudanthias ignitus) displaying to the more drab female (above) during courtship in the evening. Similan Islands, Thailand. Andaman Sea, Indian Ocean

Toque Macaque (Macaca sinica) baby peering through mothers legs, Yala National Park, Southern Province, Sri Lanka

Sumatran orangutan (Pongo abelii) female next to infant, Gunung Leuser National Park, Indonesia, UNESCO World Heritage site, November

Black korhaan (Eupodotis afra) male displaying and calling. Etosha National Park, Namibia, March

Jaguar (Panthera onca) male and female four month old cubs playing, native to Southern and Central America, captive

Jaguar (Panthera onca) mother grooming four month cub, native to Southern and Central America, captive

Jaguar (Panthera onca) female snarling, native to Southern and Central America, captive

Madagascar crested wood ibis (Lophotibis cristata) preening, endemic to dry forest of Madagascar, captive

Alewife fish (Alosa pseudoharengus), migrating up river, Northern Maine, USA. June

Sea otter (Enhydra lutris) mother and sleeping newborn pup (aged 3 days) Monterey, California, USA. *Digitally darkened scar on mothers nose

Peringueys / sidewinding adder (Bitis peringueyi) hidden in the sand, Naukluft National Park, Namibia

Grey wolf (Canis lupus) mother and two month old cubs, pups begging for food by licking mothers mouth, captive

Blue cheeked bee eater (Merops persicus) two squabbling, Oman, April

Polar bear (Ursus maritimus) spring cub sitting upright on hind legs balancing, Bernard Spit, 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species

Spotted catshark (Scyliorhinus canicula) just hatched from egg case that is on Seafan (Paramuricea clavata), Ullastres seamount, Costa Brava, Llafranch, Catalunya, Spain

Osprey (Pandion haliaetus) male and female head to head at nest, just before mating, Finland April

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